Добавлено 26-06-2010
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Число хромосом муравьёв
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Pedro LORITE & Teresa PALOMEQUE
E-mail:
plorite@ujaen.es
Dr. Pedro Lorite (contact author) & Dra. Teresa Palomeque, Departamento de Biologia Experimental, Area de Genetica, Universidad de Jaen, 23071 Jain, Spain. plorite@ujaen.es
Karyotype evolution in ants (Hymenoptera: Formicidae) with a review of the known ant chromosome numbers
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"Myrmecologische Nachrichten" ("Myrmecological News") 2010, Volume 13, Pages 89-102 (Wien, апрель 2010)
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ПРИЛОЖЕНИЕ к ОСНОВНОЙ СТАТЬЕ.
См. ниже частичный перевод статьи и список литературы.
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ЧИСЛО ХРОМОСОМ МУРАВЬЁВ
АЛФАВИТНЫЙ СПИСОК ВИДОВ
ПРИЛОЖЕНИЕ к ОСНОВНОЙ СТАТЬЕ.
Знак asterisk (*) означает, что в конце списка есть примечание по этому виду. Наибольшая вариация хромосомного набора обнаружена в примитивных подсемействах, таких как Ponerinae (n = 3 - 60) и Myrmeciinae (n = 1 - 47), в то время как у менее примитивных подсемействах хромосомные наборы менее вариабельны, как например, у Dolichoderinae (n = 5 -16), Formicinae (n = 8 -28) и Myrmicinae (n = 4 - 35).
Acantholepis (see under Lepisiota)
Acanthomyrmex (Myrmicinae)
Acanthomyrmex sp.
2n = 24, Sarawak, TJAN & al. (1986)
Acanthomyrmex sp. 1
n = 11, Malaysia, GONI & al. (1982)
Acanthomyrmex sp. 2
2n = 22, Malaysia, GONI & al. (1982)
Acanthomyrmex sp. 3
n = 11, 2n = 22, Malaysia, IMAI & al. (1983)
Acromyrmex (Myrmicinae)
Acromyrmex ambiguus
2n = 38, Uruguay, GONI & al. (1983, K)
Acromyrmex crassipinus
2n = 38, Brazil, FADINI & POMPOLO (1996, K)
Acromyrmex heyeri
2n = 38, Uruguay, GONI & al. (1983, K)
2n = 38, Brazil, SANTOS-COLARES & al. (1997, M)
Acromyrmex hispidus
2n = 38, Uruguay, GONI & al. (1983, K)
Acromyrmex molestans (see A. subterraneus)
Acromyrmex subterraneus
2n = 38, Brazil, FADINI & POMPOLO (1996, K)
Acropyga (Formicinae)
Acropyga acutiventris*
n = 15, 2n = 28-29, Malaysia, GOСI & al. (1982), IMAI & al. (1983)
Acropyga sp.
2n = 32, Indonesia, IMAI & al. (1985)
Acropyga sp.
n = 15, 2n = 30, Sarawak, TJAN & al. (1986)
Aenictus (Aenictidae)
Aenictus brevicornis
2n = 24, India, IMAI & al. (1984, K)
Aenictus laeviceps
2n = 22, Malaysia, IMAI & al. (1983)
Aenictus sp. (near A. camposi)
n = 15, 2n = 30, Taiwan, HUNG & al. (1972)
Amblyopone (Amblyoponinae)
Amblyopone australis
n = 24, 2n = 48, Australia, IMAI & al. (1977, M, K)
Amblyopone reclinata
2n = 38, Malaysia, IMAI & al. (1983)
Amblyopone cf. fortis
2n = 44, Australia, IMAI & al. (1977, K)
Anochetus (Ponerinae)
Anochetus graeffei*
2n = 30, India, IMAI & al. (1984, K)
2n = 38, Indonesia, IMAI & al. (1985)
Anochetus madaraszi
2n = 28, India, IMAI & al. (1984, K)
Anochetus modicus
2n = 30, Indonesia, IMAI & al. (1985)
Anochetus yerburyi
2n = 30, India, IMAI & al. (1984, K)
Anochetus sp. 1
2n = 24, Malaysia, GONI & al. (1982)
Anochetus sp. 2
n = 19, Malaysia, GONI & al. (1982)
Anochetus sp. 4
2n = 30, India, IMAI & al. (1984, K)
Anochetus sp. 5
2n = 34, India, IMAI & al. (1984, K)
Anochetus sp.
n = 17, 2n = 34, Sarawak, TJAN & al. (1986)
Anonychomyrma (Dolichoderinae)
Anonychomyrma itinerans
n = 8, 2n = 16, Australia, CROZIER (1968a, K, as Iridomyrmex itinerans)
Anonychomyrma sp.
2n = 16, Australia, IMAI & al. (1977, K, as Iridomyrmex sp., ANIC-10)
Anoplolepis (Formicinae)
Anoplolepis longipes (see A. gracilipes)
Anoplolepis gracilipes
n = 17, 2n = 34, Malaysia, GOСI & al. (1982), IMAI & al. (1983)
2n = 34, India, IMAI & al. (1984, K)
2n = 34, Indonesia, IMAI & al. (1985)
n = 17, Sarawak, TJAN & al. (1986)
Aphaenogaster (Myrmicinae)
Aphaenogaster beccarii*
2n = 30, Malaysia, IMAI & al. (1983)
2n = 46, India, IMAI & al. (1984, K)
Aphaenogaster depilis
2n = 34, Tunisia, HAUSCHTECK-JUNGEN & JUNGEN (1983)
Aphaenogaster famelica
n = 17, 2n = 34, Japan, IMAI & YOSIDA (1964), IMAI (1966, M), IMAI (1969), IMAI (1971, K)
Aphaenogaster fulva
2n = 36, USA, CROZIER (1977, K)
Aphaenogaster gibbosa*
n = 11, Switzerland, HAUSCHTECK-JUNGEN & JUNGEN (1983)
n = 16-17, Spain, PALOMEQUE & al. (1993b, M, K), LORITE & al. (2000, M, K)
Aphaenogaster iberica*
n = 17, 2n = 34, Spain, PALOMEQUE & al. (1993a, M, K), PALOMEQUE & al. (1993b, M, K), LORITE & al. 2000 (M, K)
Aphaenogaster lamellidens
2n = 38, USA, CROZIER (1977, K), TABER & COKEN-DOLPHER (1988)
Aphaenogaster longiceps*
2n = 45-46, Australia, IMAI & al. 1977 (K)
Aphaenogaster miamiana
2n = 36, USA, CROZIER (1977, K)
Aphaenogaster osimensis
n = 16, 2n = 32, Japan, IMAI & YOSIDA (1964, as A. sp.), IMAI & YOSIDA (1966), IMAI (1966, M), IMAI (1969), IMAI (1971, K)
Aphaenogaster rudis*
n = 16-18, 20-22, 2n = 40-42, 44, USA, CROZIER (1970b), CROZIER (1975, K), CROZIER (1977)
Aphaenogaster sardoa
2n = 34, Tunisia, HAUSCHTECK-JUNGEN & JUNGEN (1983)
Aphaenogaster senilis
n = 16, 2n = 32, Spain, PALOMEQUE & al. 1993a (M, K), PALOMEQUE & al. (1993b, M, K), LORITE & al. 2000 (M, K)
Aphaenogaster smythiesi*
n = 11, 2n = 22, Japan, IMAI (1969), IMAI (1971, K)
2n = 34, India, IMAI & al. 1984 (K)
Aphaenogaster subterranea
2n = 22, Switzerland, HAUSCHTECK (1962, M)
n = 11, 2n = 22, Switzerland and Germany, HAUSCHTECK-JUNGEN & JUNGEN (1983, K)
Aphaenogaster testaceopilosa*
n = 17, 2n = 34, Tunisia, Spain and Croatia, HAUSCHTECK-JUNGEN & JUNGEN (1983, K)
Aphaenogaster tipuna
2n = 34, Taiwan, HUNG & al. (1972)
Aphaenogaster treatae
2n = 42, USA, CROZIER (1977, K)
Aphaenogaster sp.
2n = 30, Malaysia, GOСI & al. (1982)
Apterostigma (Myrmicinae)
Apterostigma mayri
2n = 24, Panama, MURAKAMI & al. (1998, K)
Apterostigma sp.
2n = 20, Brazil, FADINI & POMPOLO (1996, K)
Apterostigma sp.
2n = 24, Panama, MURAKAMI & al. (1998, K)
Atta (Myrmicinae)
Atta bisphaerica
2n = 22, Brazil, FADINI & POMPOLO (1996, K)
Atta columbica
n = 11, 2n = 22, Panama, MURAKAMI & al. (1998, K)
Atta laevigata
2n = 22, Brazil, FADINI & POMPOLO (1996, K)
Atta sexdens
2n = 22, Brazil, FADINI & POMPOLO (1996, K), SANTOS-COLARES & al. (1997, M)
Bothriomyrmex (Dolichoderinae)
Bothriomyrmex gibbus
n = 11, Italy, HAUSCHTECK-JUNGEN & JUNGEN (1983)
Bothriomyrmex pusillus
2n = 22, Australia, IMAI & al. (1977, K)
Bothriomyrmex sp.
n = 11, Switzerland, HAUSCHTECK (1963)
Bothroponera (see under Pachycondyla)
Brachymyrmex (Formicinae)
Brachymyrmex sp.
2n = 18, Brazil, CROZIER (1970b, K)
Brachyponera (see under Pachycondyla)
Calomyrmex (Formicinae)
Calomyrmex sp. (ANIC-1)
2n = 28, Australia, IMAI & al. (1977, K)
Camponotus (Formicinae)
Camponotus aethiops
2n = 42, France, old Yugoslavia, HAUSCHTECK-JUNGEN & JUNGEN (1983, K)
Camponotus alii*
2n = 42, Spain, HAUSCHTECK-JUNGEN & JUNGEN (1983)
Camponotus atriceps
n = 20, 2n = 40, Brazil, MARIANO & al. (2001, K)
Camponotus balzani
n = 20, 2n = 40, Brazil, MARIANO & al. (2001, K)
Camponotus bonariensis
2n = 40, Uruguay, GOСI & al. (1983, K)
Camponotus caryae (see C. vitiosus)
Camponotus cingulatus
2n = 40, Brazil, MARIANO & al. (2001, K)
Camponotus compressus*
n = 10, India, KUMBKARNI (1965)
n = 20, 2n = 40, Tunisia, HAUSCHTECK-JUNGEN & JUNGEN (1983, K)
Camponotus consobrinus
n = 23, 2n = 46, Australia, IMAI & al. (1977, K)
Camponotus crassisquamis*
2n = 39-40, India, IMAI & al. (1984, K)
Camponotus crassus
n = 10, 2n = 20, Brazil, MARIANO & al. (2001, K)
Camponotus cruentatus*
n = 18, Spain, HAUSCHTECK-JUNGEN & JUNGEN (1983)
n = 20, Spain, LORITE & al. (2002b, M, K)
Camponotus dolendus
2n = 20, India, IMAI & al. (1984, K)
Camponotus festinus
n = 19, 2n = 38, Malaysia, IMAI & al. (1983)
Camponotus foreli
n = 17, 2n = 34, Spain, HAUSCHTECK-JUNGEN & JUNGEN (1983)
Camponotus japonicus*
n = 14, Japan, IMAI & YOSIDA (1964), IMAI (1966, M), (1969, as Camponotus sp.)
n = 13, 2n = 26, Japan, IMAI & KUBOTA (1972, K)
Camponotus kiusiuensis
2n = 28, Japan, IMAI & YOSIDA (1964), IMAI (1966, M) (1969, K)
Camponotus lateralis
2n = 28, Switzerland, HAUSCHTECK (1962, M)
Camponotus ligniperda
n = 14, 2n = 28, Switzerland, HAUSCHTECK (1961, M), HAUSCHTECK-JUNGEN & JUNGEN (1983)
Camponotus mitis
2n = 20, India, IMAI & al. (1984, K)
Camponotus mus
2n = 26, Uruguay, GOСI & al. (1983, K)
Camponotus obscuripes
2n = 28, Japan, IMAI (1969)
Camponotus parius
2n = 40, India, IMAI & al. (1984, K)
Camponotus pilicornis
2n = 50, Spain, HAUSCHTECK-JUNGEN & JUNGEN (1983)
Camponotus punctulatus
2n = 40, Uruguay, GOСI & al. (1983, K)
Camponotus rufipes*
n = 20, 2n = 40, Uruguay, GOСI & al. (1983, K)
2n = 39-40, Brazil, MARIANO & al. (2001, K)
Camponotus rufoglaucus*
n = 18, Tunisia, Spain, HAUSCHTECK-JUNGEN & JUNGEN (1983)
Camponotus sericeiventris
2n = 40, Brazil, MARIANO & al. (2001)
Camponotus sericeus
2n = 44, India, IMAI & al. (1984, K)
Camponotus sylvaticus
n = 20, 2n = 40, Greece, Spain, HAUSCHTECK-JUNGEN & JUNGEN (1983)
Camponotus taylori
2n = 24, India, IMAI & al. (1984, K)
Camponotus thraso
2n = 40, India, IMAI & al. (1984, K)
Camponotus tokyoensis (see C. vitiosus)
Camponotus vagus
2n = 28, Switzerland, HAUSCHTECK (1961, M)
Camponotus variegatus
2n = 26, India, IMAI & al. (1984, K)
Camponotus vitiosus (C. caryae, C. tokyoensis)
n = 9, 2n = 18, Japan, IMAI (1966a, M, as Campo-notus sp.), IMAI (1969, K, as C. caryae group sp.), IMAI & KUBOTA (1972, K)
Camponotus sp.
n = 9, 2n = 18, Japan, IMAI & YOSIDA (1964), IMAI (1966, M)
Camponotus sp. (C. impressus group)
n = 26, 2n = 52, USA, CROZIER (1970b, K)
Camponotus sp. (C. variegatus complex)
2n = 20, Taiwan, HUNG & al. (1972)
Camponotus (Myrmamblys) sp. 1
n = 9, 2n = 18, Japan, IMAI & KUBOTA (1972, K)
Camponotus sp. (ANIC-1)
n = 23, 2n = 46, Australia, IMAI & al. (1977, K)
Camponotus sp. (ANIC-2)
n = 23, Australia, IMAI & al. (1977, K)
Camponotus sp. (ANIC-3)
2n = 48, Australia, IMAI & al. (1977, K)
Camponotus sp. (ANIC-5)
2n = 32, Australia, IMAI & al. (1977, K)
Camponotus sp. (ANIC-8)
2n = 32, Australia, IMAI & al. (1977, K)
Camponotus sp. (ANIC-9)
2n = 38, Australia, IMAI & al. (1977, K)
Camponotus sp. (ANIC-10)
2n = 46, Australia, IMAI & al. (1977, K)
Camponotus sp. (ANIC-11)
2n = 32, Australia, IMAI & al. (1977, K)
Camponotus sp. (ANIC-12)
2n = 38, Australia, IMAI & al. (1977, K)
Camponotus sp. (ANIC-13)
n = 10, 2n = 20, Australia, IMAI & al. (1977, K)
Camponotus sp. (ANIC-14)
n = 19, 2n = 38, Australia, IMAI & al. (1977, K)
Camponotus sp. 1
n = 19, Malaysia, GOСI & al. (1982)
Camponotus sp. 2
n = 20, 2n = 40, Malaysia, GOСI & al. (1982), IMAI & al. (1983)
Camponotus sp. 3
2n = 38, Malaysia, GOСI & al. (1982)
Camponotus sp. 4
n = 18, 2n = 36, Malaysia, GOСI & al. (1982)
Camponotus sp. 5
n = 20, 2n = 40, Malaysia, GOСI & al. (1982)
Camponotus sp. 6
2n = 44, Malaysia, GOСI & al. (1982)
Camponotus sp. 7
n = 20, Malaysia, IMAI & al. (1983)
Camponotus sp. 8
2n = 38-39, Malaysia, IMAI & al. (1983)
Camponotus sp. 9
2n = 40, Malaysia, IMAI & al. (1983)
Camponotus sp. 10
n = 13-14, Malaysia, IMAI & al. (1983)
Camponotus sp. 11
2n = 52, Malaysia, IMAI & al. (1983)
Camponotus sp. 12
2n = 18, Malaysia, IMAI & al. (1983)
Camponotus sp. 13
2n = 18, Malaysia, IMAI & al. (1983)
Camponotus sp. 7 (near C. variegatus)
2n = 32, India, IMAI & al. (1984, K)
Camponotus sp. 9 (near C. variegatus)
2n = 34-35, India, IMAI & al. (1984, K)
Camponotus sp. 10 (near C. infuscus)
2n = 20, India, IMAI & al. (1984, K)
Camponotus sp. 12 (near C. variegatus)
2n = 34, India, IMAI & al. (1984, K)
Camponotus sp. 1
2n = 38-39, Indonesia, IMAI & al. (1985)
Camponotus sp. 1
2n = 40, Sarawak, TJAN & al. (1986)
Camponotus sp. 2
2n = 20, Sarawak, TJAN & al. (1986)
Camponotus sp. 2
2n = 38, Indonesia, IMAI & al. (1985)
Camponotus (Myrmobrachys) sp.*
2n = 40, 41-43, Brazil, MARIANO & al. (2001, K)
Cardiocondyla (Myrmicinae)
Cardiocondyla nuda
2n = 28, India, IMAI & al. (1984, K)
Cardiocondyla sp
2n = 40, Malaysia, GOСI & al. (1982)
Carebara (Myrmicinae)
Carebara asina (Oligomyrmex asinus)
2n = 44, India, IMAI & al. (1984, K)
Carebara sauteri (Oligomyrmex sauteri)
n = 18, Taiwan, HUNG & al. (1972)
Carebara sp. (ANIC-6)
2n = 38, Australia, IMAI & al. (1977, K, as Oligomyr-mex sp., ANIC-6)
Carebara sp. 1
2n = 36, Malaysia, GOСI & al. (1982, as Oligomyrmex sp. 1)
Carebara sp. 2
2n = 44, India, IMAI & al. (1984, K, as Oligomyrmex sp. 2)
Carebara sp. 4
n = 16, 2n = 32, India, IMAI & al. (1984, K, as Oligo-myrmex sp. 4)
Carebara sp. 5
2n = 26, India, IMAI & al. (1984, K, as Oligomyrmex sp. 5)
Carebara sp. 1
2n = 34, Indonesia, IMAI & al. (1985, as Oligomyrmex sp. 1)
Carebara sp. 2
2n = 42, Indonesia, IMAI & al. (1985, as Oligomyrmex sp. 2)
Carebara sp. 1
2n = 36, Sarawak, TJAN & al. (1986, as Oligomyrmex sp. 1)
Carebara sp. 2
2n = 44, Sarawak, TJAN & al. (1986, as Oligomyrmex sp. 2)
Cataglyphis (Formicinae)
Cataglyphis albicans (see C. iberica)
Cataglyphis bicolor
n = 26, Israel, Tunisia, HAUSCHTECK-JUNGEN & JUN-GEN (1983)
Cataglyphis iberica*
n = 26, Spain, HAUSCHTECK-JUNGEN & JUNGEN (1983, as C. albicans)
Cataglyphis setipes
2n = 54, India, IMAI & al. (1984, K)
Centromyrmex (Ponerinae)
Centromyrmex feae
2n = 44, India, IMAI & al. (1984, K)
Cerapachys (Cerapachyinae)
Cerapachys biroi
2n = 28, India, IMAI & al. (1984, K)
Cerapachys brevis
n = 23, 2n = 46, Australia, IMAI & al. (1977, K)
Cerapachys sp.
2n = 50, Sarawak, TJAN & al. (1986)
Cerapachys sp.
n = 25, 2n = 50, Malaysia, GOСI & al. (1982)
Chalepoxenus (Myrmicinae)
Chalepoxenus kutteri
n = 12, France, FISCHER (1987, K)
Chalepoxenus muellerianus (C. siciliensis)
n = 12, Italy, France, Croatia, Greece, FISCHER (1987, K)
Chelaner (see under Monomorium)
Colobostruma (Myrmicinae)
Colobostruma alinodis
n = 11, 2n = 22, Australia, CROZIER (1968d, M)
Colobostruma sp.
n = 10, 2n = 20, Australia, CROZIER (1968d, M, as Epopostruma sp.)
Colobostruma sp. (ANIC-1)
2n = 22, Australia, IMAI & al. (1977, K)
Conomyrma (see under Dorymyrmex)
Crematogaster
Crematogaster biroi
2n = 24, India, IMAI & al. (1984, K)
Crematogaster brunnea
2n = 36, India, IMAI & al. (1984, K)
Crematogaster laboriosa
2n = 26, Japan, IMAI & YOSIDA (1964), IMAI (1966, M), IMAI (1969, K)
Crematogaster rothneyi
2n = 50, India, IMAI & al. (1984, K)
Crematogaster subnuda
n = 18, 2n = 36, India, IMAI & al. (1984, K)
Crematogaster sp.
2n = 40, Japan, IMAI (1969, K)
Crematogaster sp. (ANIC-1)
n = 12, 2n = 24, Australia, IMAI & al. (1977, K)
Crematogaster sp. (ANIC-2)
2n = 26, 39, Australia, IMAI & al. (1977, K)
Crematogaster sp. 1
2n = 26, Malaysia, GOСI & al. (1982), IMAI & al. (1983)
Crematogaster sp. 2
2n = 36, Malaysia, GOСI & al. (1982)
Crematogaster sp. 3
2n = 36, Malaysia, GOСI & al. (1982)
Crematogaster sp. 4
2n = 24, Malaysia, GOСI & al. (1982), IMAI & al. (1983)
Crematogaster sp. 5
2n = 26, Malaysia, IMAI & al. (1983)
Crematogaster sp. 6
2n = 26, Malaysia, IMAI & al. (1983)
Crematogaster sp. 1
2n = 24, Indonesia, IMAI & al. (1985)
Crematogaster sp. 2
2n = 56, 58, Indonesia, IMAI & al. (1985)
Crematogaster sp.
2n = 26, Sarawak, TJAN & al. (1986)
Cryptopone (Ponerinae)
Cryptopone rotundiceps
2n = 12, Australia, IMAI & al. (1977, K)
Cryptopone sauteri
n = 14, 2n = 28, Japan, IMAI & YOSIDA (1964), IMAI (1969, K), IMAI & KUBOTA (1972, K)
Cryptopone testacea
n = 9, 2n = 18, Malaysia, IMAI & al. (1983)
n = 9, 2n = 18, Sarawak, TJAN & al. (1986)
Cylindromyrmex (Cerapachyinae)
Cylindromyrmex brasiliensis
2n = 34, Brazil, MARIANO & al. (2004b, M, K)
Cyphomyrmex (Myrmicinae)
Cyphomyrmex costatus
2n = 20, Panama, MURAKAMI & al. (1998, K)
Cyphomyrmex rimosus
2n = 32, Panama, MURAKAMI & al. (1998, K)
Dacetinops (Myrmicinae)
Dacetinops concinnus
2n = 16, Malaysia, IMAI & al. (1983)
Diacamma (Ponerinae)
Diacamma rugosum (D. vagans)
n = 7, 2n = 14, India, IMAI & al. (1984, K)
Diacamma vagans (see D. rugosum)
Diacamma sp.
n = 18, 2n = 36, Malaysia, GOСI & al. (1982)
Diacamma sp. 1
2n = 44, Malaysia, IMAI & al. (1983)
Diacamma sp. 2
2n = 30, India, IMAI & al. (1984)
Diacamma sp.
2n = 66, Indonesia, IMAI & al. (1985)
Diacamma sp.
2n = 58, Sarawak, TJAN & al. (1986)
Diceratoclinea (see under Dolichoderus)
Dinoponera (Ponerinae)
Dinoponera lucida*
n = 57-60, 2n = 106, 116, 118, 120, Brazil, MARIANO & al. (2004a, M), MARIANO & al. (2008, M)
Diplorhoptrum (see under Solenopsis)
Discothyrea (Proceratiinae)
Discothyrea sp. (near D. bryanti)
2n = 30, Indonesia, IMAI & al. (1985)
Doleromyrma (Dolichoderinae)
Doleromyrma sp. ("darwinianus-group")
n = 7, 2n = 14, Australia, IMAI & al. (1977, K, as Iri-domyrmex sp., ANIC-8)
Doleromyrma sp. ("darwinianus-group")
2n = 12, Australia, IMAI & al. (1977, K, as Iridomyrmex sp., ANIC-9)
Dolichoderus (Dolichoderinae)
Dolichoderus bituberculatus (see D. thoracicus)
Dolichoderus quadripunctatus
2n = 28, Japan, IMAI (1969)
Dolichoderus scabridus (Diceratoclinea scrabida)
n = 14, 2n = 28, Australia, CROZIER (1966), IMAI & al. (1977, M)
Dolichoderus thoracicus (D. bituberculatus)*
2n = 30-33, Malaysia, IMAI & al. (1983)
2n = 30, Indonesia, IMAI & al. (1985)
Dolichoderus sp.
2n = 18, Malaysia, GOСI & al. (1982)
Doronomyrmex (see under Leptothorax)
Dorymyrmex (Dolichoderinae)
Dorymyrmex bicolor
n = 13, 2n = 26, USA, CROZIER (1970b, K)
Dorymyrmex flavus (Conomyrma flava)
2n = 26, USA, COKENDOLPHER & FRANCKE (1984, K)
Dorymyrmex pulchellus
2n = 18, Brazil, CROZIER (1968b, M, as Dorymyrex sp.) (1970b, K)
Dorymyrmex pyramicus
2n = 18, Uruguay, GOСI & al. (1983, K)
Dorymyrmex thoracicus
n = 9, Peru, CROZIER (1970b, K)
Echinopla (Formicinae)
Echinopla sp. 1
n = 12, 2n = 24, Malaysia, IMAI & al. (1983)
Ectatomma (Ectatomminae)
Ectatomma brunneum
2n = 44, Brazil, BARROS & al. (2008, K)
Ectatomma edentatum
2n = 46, Brazil, BARROS & al. (2008)
Ectatomma muticum
n = 20, Brazil, BARROS & al. (2008, K)
Ectatomma permagnum
2n = 46, Brazil, BARROS & al. (2008, K)
Ectatomma tuberculatum
2n = 36, Brazil, BARROS & al. (2008, K)
Ectomomyrmex (see under Pachycondyla)
Epimyrma (see under Myrmoxenus)
Epopostruma (see under Colobostruma)
Eurhopalothrix (Myrmicinae)
Eurhopalothrix sp. (E. procera group)
2n = 18, Malaysia, IMAI & al. (1983)
Forelius (Dolichoderinae)
Forelius foetida (see F. mccooki)
Forelius mccooki (F. foetida)
n = 16, 2n = 32, USA, CROZIER (1970b, K), CROZIER (1975, K)
Formica (Formicinae)
Formica aquilonia
n = 26, Finland, ROSENGREN & al. (1980, M)
Formica candida (F. picea, F. transkaucasica)
2n = 52, Japan, IMAI (1969)
n = 26, 2n = 52, Switzerland, HAUSCHTECK-JUNGEN & JUNGEN (1976, M),
n = 26, Finland, ROSENGREN & al. (1980)
Formica cinerea
n = 27, 2n = 54, Switzerland, HAUSCHTECK-JUNGEN & JUNGEN (1976, K)
Formica cunicularia
n = 27, 2n = 54, France, HAUSCHTECK-JUNGEN & JUNGEN (1976, K)
Formica dakotensis
n = 26, HUNG (in CROZIER 1975)
Formica exsecta
n = 26, 2n = 52, Switzerland, AGOSTI & HAUSCHTECK-JUNGEN (1987), HAUSCHTECK-JUNGEN & JUNGEN (1976, M)
Formica frontalis
2n = 52, Spain, LORITE & al. (2002b, M, K), LORITE & al. (2004a, M)
Formica fusca
2n = 54, Japan, IMAI (1969)
n = 27, Germany, Switzerland, HAUSCHTECK-JUNGEN & JUNGEN (1976, M)
Formica gagates
n = 27, 2n = 54, France, old Yugoslavia, HAUSCHTECK-JUNGEN & JUNGEN (1976)
Formica gerardi
n = 27, Spain, LORITE & al. (1998a, M, K) (2002b, M, K)
Formica japonica
n = 27, 2n = 54, Japan, IMAI & YOSIDA (1964), IMAI (1966, M), IMAI (1969), CROZIER (1975, M)
Formica lemani
2n = 54, Switzerland, HAUSCHTECK-JUNGEN & JUNGEN (1976)
Formica lugubris (F. nylanderi)
n = 26, 2n = 52, Switzerland, HAUSCHTECK-JUNGEN & JUNGEN (1976, M)
n = 26, Finland, ROSENGREN & al. (1980, M)
Formica montana
n = 27, 2n = 54, USA, HUNG (1969, M)
Formica nylanderi (see F. lugubris)
Formica obscuripes
n = 26, USA, HUNG (1969, M)
Formica pallitarsis (see Lasius pallitarsis)
Formica picea (see F. candida)
Formica pergandei
n = 26, HUNG in CROZIER (1975)
Formica polyctena
n = 26, 2n = 52, Switzerland, HAUSCHTECK-JUNGEN & JUNGEN (1976, M)
n = 26, Finland, ROSENGREN & al. (1980)
Formica pratensis
n = 26, 2n = 52, Switzerland, HAUSCHTECK-JUNGEN & JUNGEN (1976, M)
n = 26, Finland, ROSENGREN & al. (1980)
Formica pressilabris
n = 26, Finland, ROSENGREN & al. (1980)
Formica reflexa
n = 26, HUNG in CROZIER (1975)
Formica rufa
n = 26, 2n = 52, Switzerland, HAUSCHTECK-JUNGEN & JUNGEN (1976, M)
Formica rufibarbis
n = 27, 2n = 54, Switzerland, HAUSCHTECK-JUNGEN & JUNGEN (1976, K)
n = 27, Finland, ROSENGREN & al. (1980)
Formica sanguinea
n = 26, 2n = 52, Japan, IMAI & YOSIDA (1964), IMAI (1966, M), IMAI (1969, K)
n = 26, 2n = 52, Switzerland, HAUSCHTECK-JUNGEN & JUNGEN (1976, M)
Formica subintegra
n = 26, USA, HUNG (1969)
Formica subrufa
n = 26, Spain, LORITE & al. (2002b, M, K)
Formica transkaucasica (see F. candida)
Formica truncorum*
n = 26, 2n = 52, Japan, IMAI & YOSIDA (1964), IMAI (1969)
n = 26, 28, 2n = 52, Switzerland, HAUSCHTECK-JUNGEN & JUNGEN (1976, K)
n = 26-28, Finland, ROSENGREN & al. (1980, M)
Formica ulkei
n = 26, USA, HUNG (1969, M).
Formica uralensis
n = 26, Finland, ROSENGREN & al. (1980, M)
Formica yessensis
n = 26, 2n = 52, Japan, IMAI (1966, M), (1969)
Formica 3 spp. (F. fusca group)
n = 27, HUNG (in CROZIER 1975)
Formica 4 spp.
n = 26, HUNG (in CROZIER 1975)
Formicoxenus (Myrmicinae)
Formicoxenus chamberlini (Symmyrmica chamberlini)
2n = 28, USA, BUSCHINGER & FRANCOEUR (1983), FRANCOEUR & al. (1985), FISCHER (1987, M)
Formicoxenus hirticolis*
2n = 30-34, FRANCOEUR & al. (1985)
Formicoxenus nitidulus
n = 15, France, BUSCHINGER & al. (1980), FRANCOEUR & al. (1985), FISCHER (1987, M)
Formicoxenus provancheri (Leptothorax provancheri)
n = 11, 2n = 22, Canada, BUSCHINGER & al. (1980, K), FISCHER (1987, K)
Formicoxenus quebecensis
2n = 28, Canada, FRANCOEUR & al. (1985, M), FISCHER (1987, M)
Gnamptogenys (Ectatomminae)
Gnamptogenys annulata
2n = 68, Brazil, BORGES & al. (2004b, K)
Gnamptogenys binghamii
n = 22, Malaysia, IMAI & al. (1983)
Gnamptogenys menadensis
2n = 42, Malaysia, GOСI & al. (1982, as G. sp. 1), IMAI & al. (1983)
Gnamptogenys striatula
2n = 34, Brazil, BORGES & al. (2004b, K)
Gnamptogenys sp. 2
2n = 36, Malaysia, GOСI & al. (1982)
Gnamptogenys sp.
n = 23, 2n = 46, Brazil, BORGES & al. (2004b, K)
Harpagoxenus (Myrmicinae)
Harpagoxenus americanus (see Protomognathus americanus)
Harpagoxenus canadensis
n = 18, Canada, BUSCHINGER & al. (1980), FISCHER (1987, K)
Harpagoxenus sublaevis
n = 20, BUSCHINGER & al. (1980), FISCHER (1987, K)
n = 20, 2n = 40, Switzerland, HAUSCHTECK-JUNGEN & JUNGEN (1983, M)
Heteroponera (Heteroponerinae)
Heteroponera dolo
n = 12, 2n = 24, Brazil, BORGES & al. (2004a)
Heteroponera relicta
n = 11, 2n = 22, Australia, IMAI & al. (1977, K)
Hypoponera (Ponerinae)
Hypoponera confinis
2n = 38, Indonesia, IMAI & al. (1985)
Hypoponera pruinosa
n = 12, 2n = 24, Malaysia, IMAI & al. (1983)
2n = 24, Indonesia, IMAI & al. (1985)
Hypoponera sp. (ANIC-1)
n = 19, 2n = 38, Australia, IMAI & al. (1977, K)
Hypoponera sp. (ANIC-2)
2n = 38, Australia, IMAI & al. (1977, K)
Hypoponera sp.
2n = 38, Malaysia, GOСI & al. (1982)
Hypoponera sp. 2
2n = 38, Malaysia, IMAI & al. (1983)
Hypoponera sp. 3
2n = 36, Malaysia, IMAI & al. (1983)
Iridomyrmex (Dolichoderinae)
Iridomyrmex anceps*
2n = 18, India, IMAI & al. (1984, K)
2n = 48, Indonesia, IMAI & al. (1985)
Iridomyrmex cordata (see Philidris cordata)
Iridomyrmex detecta (see I. purpureus)
Iridomyrmex glaber (see Ochetellus glaber)
Iridomyrmex gracilis
n = 9, 2n = 18, Australia, CROZIER (1968a, K)
Iridomyrmex humilis (see Linepithema humile)
Iridomyrmex itinerans (see Anonychomyrma itinerans)
Iridomyrmex itoi (see Ochetellus glaber)
Iridomyrmex mattiroloi
n = 9, 2n = 18, Australia, CROZIER (1968a, K, M)
Iridomyrmex nitidus (see Papyrius nitidus)
Iridomyrmex pilifer (see Linepithema pilifer)
Iridomyrmex purpureus (I. detectus)*
n = 9, 2n = 18, Australia, CROZIER (1968a, K)(1968c, M, K) IMAI & al. (1977, K)
Iridomyrmex sp. (ANIC-5)*
n = 7, 2n = 14, Australia, CROZIER (1968a, K)
Iridomyrmex sp. (ANIC-6)
n = 9, 2n = 18, Australia, CROZIER (1968a, K)
Iridomyrmex sp. (ANIC-11)
n = 9, 2 = 18, Australia, CROZIER (1968a, K)
Iridomyrmex sp. (ANIC-12)
2n = 18, Australia, CROZIER (1968a, K)
Iridomyrmex sp. (ANIC-13)
2n = 18, Australia, IMAI & al. (1977, M, K)
Iridomyrmex sp. (ANIC-14)
2n = 18, Australia, IMAI & al. (1977, K)
Iridomyrmex sp. (ANIC-15)
2n = 18, Australia, IMAI & al. (1977, K)
Iridomyrmex sp. (ANIC-16)
2n = 18, Australia, IMAI & al. (1977, K)
Iridomyrmex sp. (ANIC-17)
2n = 18, Australia, IMAI & al. (1977, K)
Iridomyrmex sp.
2n = 18, Malaysia, GOСI & al. (1982)
Kyidris (see under Pyramica)
Lasius (Formicinae)
Lasius alienus*
2n = 28, Switzerland, HAUSCHTECK (1962, M)
2n = 30, Switzerland, Germany, Spain, HAUSCHTECK-JUNGEN & JUNGEN (1983, K)
Lasius brunneus*
n = 15, Spain, LORITE & al. (1998a M, K), LORITE & al. (2002b)
Lasius emarginatus
2n = 30, Switzerland, HAUSCHTECK (1962, M)
Lasius flavus
n = 15, 2n = 30, Switzerland, HAUSCHTECK (1962, M), HAUSCHTECK-JUNGEN & JUNGEN (1983, K)
Lasius fuliginosus
n = 14, 2n = 28, Switzerland, Italy, HAUSCHTECK (1962, M), HAUSCHTECK-JUNGEN & JUNGEN (1983)
Lasius nearcticus
2n = 30, USA, CROZIER (1970b, K)
Lasius niger*
2n = 30, Switzerland, HAUSCHTECK (1962, M), HAUSCHTECK (1963)
n = 15, 2n = 30, Japan, IMAI & YOSIDA (1964), IMAI (1966a, M), IMAI (1969, K), IMAI & KUBOTA (1972, K)
n = 15, 2n = 30, Spain, PALOMEQUE & al. (1990b, M, K), PALOMEQUE & al. (1990c, M, K)
Lasius pallitarsus
2n = 28, USA, HUNG (1969, M)
Lasius sakagamii
n = 15, Japan, YAMAUCHI & al. (2001, M)
Lasius talpa
2n = 30, Japan, IMAI & YOSIDA (1964), IMAI (1966, M), IMAI (1969)
Lasius umbratus
n = 15, USA, HUNG (1969, M)
2n = 30, Switzerland, HAUSCHTECK (1962, M)
Lepisiota (Formicinae)
Lepisiota capensis (L. lunaria, Acantholepis lunaria, A. capensis)
2n = 18, India, IMAI & al. (1984, K)
Lepisiota sp. 1 (near L. sericea)
2n = 18, India, IMAI & al. (1984, K, as Acantholepis sp. 1, near sericea)
Lepisiota sp. 2 (near L. fergusoni)
2n = 18, India, IMAI & al. (1984, K, as Acantholepis sp. 2, near fergusoni)
Leptogenys (Ponerinae)
Leptogenys borneensis
2n = 46, Malaysia, IMAI & al. (1983)
Leptogenys diminuta*
2n = 38, Malaysia, GOСI & al. (1982, as L. sp. 1), IMAI & al. (1983)
2n = 38, India, IMAI & al. (1984, K)
2n = 32, Indonesia, IMAI & al. (1985)
Leptogenys hysterica
2n = 26, India, IMAI & al. 1984, K)
Leptogenys iridescens
2n = 46, Malaysia, IMAI & al. (1983)
2n = 46, Indonesia, IMAI & al. (1985)
Leptogenys kraepelini
2n = 26, Indonesia, IMAI & al. (1985)
Leptogenys minchinii
2n = 52, India, IMAI & al. 1984, K)
Leptogenys myops
n = 24, 2n = 48, Indonesia, IMAI & al. (1983), IMAI & al. (1985)
2n = 48, Malaysia, GOСI & al. (1982, as L. sp. 2)
2n = 48, Sarawak, TJAN & al. (1986)
Leptogenys ocellifera (see L. processionalis)
Leptogenys peugueti
2n = 54, Indonesia, IMAI & al. (1985)
Leptogenys processionalis (L. ocellifera)
2n = 46, India, IMAI & al. (1984, K)
Leptogenys sp. 3
2n = 48, Malaysia, IMAI & al. (1983)
Leptogenys sp. 5 (near L. peugueti)
n = 15, 2n = 30, India, IMAI & al. (1984, K)
Leptogenys sp.
2n = 54, Sarawak, TJAN & al. (1986)
Leptomyrmex (Dolichoderinae)
Leptomyrmex erytrocephalus
n = 12 Australia, IMAI & al. (1977, K)
Leptothorax (Myrmicinae)
Leptothorax acervorum*
n = 13, 2n = 26, Germany, Switzerland, HAUSCH-TECK-JUNGEN & JUNGEN (1983, K)
n = 13, Germany, Switzerland, Sweden, France, Canada, FISCHER (1987, K)
n = 13, Canada, LOISELLE & al. (1990)
Leptothorax affinis (see Temnothorax affinis)
Leptothorax albipennis (see Temnothorax albipennis)
n = 8, Great Britain, ORLEDGE (1998, M, K)
n = 8, Germany, FISCHER (1987, K)
Leptothorax ambiguus (see Temnothorax ambiguus)
Leptothorax andrei (see Temnothorax andrei)
Leptothorax angustulus (see Temnothorax angustulus)
Leptothorax congruus (see Temnothorax congruus)
Leptothorax corticalis (see Temnothorax corticalis)
Leptothorax crassipilis*
n = 17-18, USA, FRANCOEUR (1986), LOISELLE & al. (1990, M)
Leptothorax curvispinosus (see Temnothorax curvispinosus)
Leptothorax duloticus (see Temnothorax duloticus)
Leptothorax exilis (see Temnothorax exilis)
Leptothorax faberi
n = 15, Canada, BUSCHINGER (1982, K)
n = 15-16, Canada, FISCHER (1987, K)
Leptothorax flavicornis (see Temnothorax flavicornis)
Leptothorax goesswaldi (Doronomyrmex goesswaldi)
n = 28, Switzerland, France, FISCHER (1987, M)
Leptothorax gredleri
n = 11, Switzerland, Germany, BUSCHINGER & al. (1980), FISCHER (1987, K), LOISELLE & al. (1990)
Leptothorax gredosi (seeTemnothorax gredosi)
Leptothorax interruptus (see Temnothorax interruptus)
Leptothorax kraussei (see Temnothorax kraussei)
Leptothorax kutteri (Doronomyrmex kutteri)*
n = 23, Sweden, DOUWES & BUSCHINGER (1983, M)
n = 23, 24-25, Sweden, Germany, Switzerland, BUSCHINGER & FISCHER (1991, M)
n = 23, 25, Sweden, Germany, Switzerland, France, Italy, FISCHER (1987, M)
Leptothorax lichtensteini (see Temnothorax lichtensteini)
Leptothorax longispinosus (see Temnothorax longispinosus)
Leptothorax melas (see Temnothorax melas)
Leptothorax muscorum*
n = 17, BUSCHINGER & al. (1980)
n = 17-18, Germany, Canada, USA, FISCHER (1987, K)
n = 16-18, 22-23, 2n = 34-36, 44, Canada, USA, LOISELLE & al. (1990, M)
Leptothorax niger (see Temnothorax niger)
Leptothorax nigriceps (see Temnothorax nigriceps)
Leptothorax nylanderi (see Temnothorax nylanderi)
Leptothorax pacis (Doronomyrmex pacis)
n = 26, Switzerland, France, FISCHER (1987, M), BUSCHINGER & FISCHER (1991)
Leptothorax parvulus (see Temnothorax parvulus)
Leptothorax pocahontas (Doronomyrmex pocahontas)
n = 18, Canada, FISCHER (1987, K), BUSCHINGER & HEINZE (1993)
Leptothorax provancheri (see Formicoxenus provancheri)
Leptothorax rabaudi (see Temnothorax rabaudi)
Leptothorax racovitzai (see Temnothorax racovitzai)
Leptothorax recedens (see Temnothorax recedens)
Leptothorax retractus*
n = 17-18, Canada, FRANCOEUR (1986), LOISELLE & al. (1990, M)
Leptothorax rottenbergii (see Temnothorax rottenbergii)
Leptothorax rougeti (see Temnothorax unifasciatus)
Leptothorax rugatulus (see Temnothorax rugatulus)
Leptothorax schaumii (seeTemnothorax schaumii)
Leptothorax sordidulus (see Temnothorax sordidulus)
Leptothorax specularis (see Temnothorax specularis)
Leptothorax sphagnicola
n = 13, Canada, FRANCOEUR (1986), FISCHER (1987, K), LOISELLE & al. (1990)
Leptothorax spinosior (see Temnothorax spinosior)
Leptothorax spinosus (see Temnothorax spinosus)
Leptothorax tristis (see Temnothorax tristis)
Leptothorax tuberum (seeTemnothorax tuberum)
Leptothorax tuberointerruptus (see Temnothorax albipennis)
Leptothorax unifasciatus (see Temnothorax unifasciatus)
Leptothorax sp.
n = 16, Japan, IMAI (1969)
Leptothorax spp.
n = 21, France, FISCHER (1987, K)
Leptothorax sp. 1
n = 17, USA, FISCHER (1987)
Leptothorax sp. 2
n = 12, Algeria, FISCHER (1987, K)
Leptothorax sp. 3
n = 12, Italy, FISCHER (1987, K)
Leptothorax sp. 4
n = 13, Croatia, FISCHER (1987, K)
Leptothorax sp. 5
n = 14, Canada, FISCHER (1987, K)
Leptothorax sp. 6
n = 22, France, FISCHER (1987)
Leptothorax sp. 1
n = 15-16, Canada, LOISELLE & al. (1990, M)
Leptothorax sp. A
n = 17, USA, LOISELLE & al. (1990, M)
Linepithema (Dolichoderinae)
Linepithema humile (Iridomyrmex humilis)
n = 8, 2n = 16, Australia, CROZIER (1968a, K), CRO-ZIER (1975, K)
n = 8, 2n = 16, Spain, LORITE & al. (1996b, M, K), LORITE & al. (1998b, M, K)
Linepithema pilifer (Iridomyrmex pilifer)
2n = 18, Peru, CROZIER (1970b, K)
Linepithema sp.
2n = 18, Peru, CROZIER (1970b, K, as Iridomyrmex sp. nr. pilifer)
Lophomyrmex (Myrmicinae)
Lophomyrmex bedoti
2n = 38, Malaysia, IMAI & al. (1983)
2n = 38, India, IMAI & al. (1984)
Lophomyrmex sp.
2n = 38, Sarawak, TJAN & al. (1986)
Lordomyrma (Myrmicinae)
Lordomyrma sp. 1
n = 11, 2n = 22, Malaysia, IMAI & al. (1983)
Manica (Myrmicinae)
Manica rubida (Myrmica rubida)
2n = 44, Switzerland, HAUSCHTECK (1965), HAUSCH-TECK-JUNGEN & JUNGEN (1983, M)
Mayriella (Myrmicinae)
Mayriella abstinens
2n = 18, Australia, IMAI & al. (1977, K)
Meranoplus (Myrmicinae)
Meranoplus bicolor
2n = 16, India, IMAI & al. (1984, K),
2n = 16, Indonesia, IMAI & al. (1985)
Meranoplus minor
2n = 22, Australia, IMAI & al. (1977, K)
Meranoplus sp. (M. oceanicus group)
n = 10, Australia, CROZIER (1966)
Meranoplus sp. (M. hirsutus group)
2n = 22, Australia, CROZIER (1970b, K)
Meranoplus sp. (ANIC-4)
2n = 22, Australia, IMAI & al. (1977, K)
Meranoplus sp. (ANIC-5)
2n = 22, Australia, IMAI & al. (1977, K)
Mesoponera (see under Pachycondyla)
Messor (Myrmicinae)
Messor aciculatus
n = 22, 2n = 44, Japan, IMAI & YOSIDA (1964), IMAI (1966a, M), IMAI (1969, K)
Messor andrei (Veromessor andrei)
2n = 40, (USA), TABER & COKENDOLPHER (1988, K)
Messor barbarus
n = 21, HAUSCHTECK-JUNGEN (In CROZIER 1975)
n = 21, Spain, LORITE & al. (2002, M, K)
Messor sp.
2n = 41, India, IMAI & al. (1984, K)
Monomorium (Myrmicinae)
Monomorium dichroum
2n = 16, India, IMAI & al. (1984, K)
Monomorium glabrum
2n = 38, India, IMAI & al. (1984, K)
Monomorium indicum*
2n = 21-22, India, IMAI & al. (1984, K)
Monomorium latinode
2n = 70, India, IMAI & al. (1984, K)
Monomorium minimum
2n = 22, USA, CROZIER (1970b, K)
Monomorium orientale
2n = 20, India, IMAI & al. (1984, K)
Monomorium pharaonis
n = 11, 2n = 22, USA, SMITH & PEACOCK (1957, M)
n = 11, Japan, IMAI & YOSIDA (1964), IMAI (1966, M), IMAI (1969, K)
Monomorium rothsteini (Chelaner rothsteini)
2n = 22, Australia, IMAI & al. (1977, K)
Monomorium scabriceps
2n = 38, India, IMAI & al. (1984, K)
Monomorium subopacum
2n = 34, Spain, LORITE & al. (2002b, M, K), LORITE & al. (2004b, M)
Monomorium viride
2n = 22, USA, CROZIER (1970b, K)
Monomorium whitei (Chelaner whitei)
2n = 24, Australia, IMAI & al. (1977, K)
Monomorium sp. (ANIC-1)
2n = 22, Australia, IMAI & al. (1977, K)
Monomorium sp. (ANIC-2)
2n = 42, Australia, IMAI & al. (1977, K)
Monomorium sp.
2n = 32, Australia, IMAI & al. (1977, K, as Chelaner sp., ANIC-1)
Monomorium sp.
2n = 22, Australia, IMAI & al. (1977, K, as Chelaner sp., ANIC-2)
Monomorium sp. 1
2n = 22, Malaysia, GOСI & al. (1982)
Monomorium sp. 2
n = 11, 2n = 22, Malaysia, GOСI & al. (1982), IMAI & al. (1983)
Monomorium sp. 3
n = 11, 2n = 22, Malaysia, IMAI & al. (1983)
Monomorium sp. 4
n = 11, Malaysia, IMAI & al. (1983)
Monomorium sp. 5
2n = 34, India, IMAI & al. (1984, K)
Monomorium sp. 6 (near M. glabrum)
2n = 38, India, IMAI & al. (1984, K)
Monomorium sp. 1
2n = 22, Indonesia, IMAI & al. (1985)
Monomorium sp. 2
2n = 22, Indonesia, IMAI & al. (1985)
Mycocepurus (Myrmicinae)
Mycocepurus sp.
2n = 8, Panama, MURAKAMI & al. (1998, K)
Myrmecia (Myrmeciinae)
Myrmecia banksi*
2n = 9-10, Australia, IMAI & al. (1994, K).
2n = 10, Australia, HIRAI & al. (1994, M, K), HIRAI & al. (1996, M)
Myrmecia brevinoda
2n = 84, Australia, IMAI & al. (1977, M. K)
Myrmecia cephalotes
2n = 66, Australia, IMAI & al. (1977, K)
Myrmecia chasei
2n = 47, Australia, MEYNE & al. (1995, M), HIRAI & al. (1996)
Myrmecia croslandi*
n = 1-2, 2n = 2-4, Australia, Taylor (1991), IMAI & al. (1992, M, 1994, K), MEYNE & al. (1995, M), HIRAI & al. (1994, M, K), HIRAI & al. (1996, M)
Myrmecia forficata*
2n = 50-52, 54, Australia, IMAI & al. (1977, K), MEYNE & al. (1995), HIRAI & al. (1996, M)
Myrmecia fulvipes*
n = 6, 2n = 12, 48, 50, 60, Australia, IMAI & al. (1977, K), MEYNE & al. (1995), HIRAI & al. (1996)
Myrmecia gulosa
2n = 38 Australia, IMAI & al. (1977, K), MEYNE & al. (1995, M), HIRAI & al. (1996, M)
Myrmecia haskinsorum*
2n = 12-24, Australia, IMAI & al. (1994, K), MEYNE & al. (1995, M), HIRAI & al. (1996, M)
Myrmecia imaii*
2n = 6-8, Australia, IMAI & al. (1994, K), HIRAI & al. (1994, M, K), HIRAI & al. (1996, M)
Myrmecia mandibularis*
n = 28-29, 2n = 56, Australia, HIRAI & al. (1996, M)
Myrmecia michaelseni
2n = 27, Australia, HIRAI & al. (1996)
Myrmecia nigrocincta
2n = 22, Australia, IMAI & al. (1977, K)
Myrmecia occidentalis
2n = 64, Australia, HIRAI & al. (1996, M)
Myrmecia pavida
2n = 44, Australia, HIRAI & al. (1996, M)
Myrmecia piliventris*
n = 2-4, 34, 2n = 4, 6, 34, 64, Australia, IMAI & TAYLOR (1986, K), IMAI & al. (1988a, K)
Myrmecia pilosula (complex) (M. ruginoda, Ponera rugi-nosa)*
n = 15, Australia, CROZIER (1966)
2n = 9-10, 30-31, Australia, IMAI & al. (1977, M, K)
n = 1, 2n = 2, Australia, CROSLAND & CROZIER (1986, M)
2n = 2, 9-10, 18-27, 30-32, Australia, CROSLAND & al. (1988)
n = 1, 2n = 2, 24-26, 31-32, Australia, IMAI & al. (1988a, K)
2n = 2-4, Australia, IMAI & al. (1988b, K)
2n = 2-4, Australia, IMAI & TAYLOR (1989, M, K)
Myrmecia pilosula (s.st.)*
2n = 18-32, Australia, IMAI & al. (1994, K), MEYNE & al. (1995, M), HIRAI & al. (1994, M, K), HIRAI & al. (1996, M)
Myrmecia pyriformis*
n = 41, 2n = 81, Australia, IMAI & al. (1977, K)
Myrmecia ruginoda (see M. pilosula)
Myrmecia simillima
2n = 70, Australia, HIRAI & al. (1996, M)
Myrmecia tepperi
2n = 70, BROWING (1987, in IMAI & al. 1990).
Myrmecia vindex*
2n = 74, 76, Australia, HIRAI & al. (1996)
Myrmecia cf. arnoldi
2n = 53, 55, 57, 59-60, 64, 66, Australia, MEYNE & al. (1995), HIRAI & al. (1996, M)
Myrmecia cf. fulvipes
2n = 12, Australia, IMAI & al. (1977)
Myrmecina (Myrmicinae)
Myrmecina americana
n = 14, HAUSCHTECK-JUNGEN (In CROZIER, 1975)
Myrmecina graminicola
n = 14, HAUSCHTECK-JUNGEN (In CROZIER 1975)
Myrmecina sp. 1
2n = 66, Indonesia, IMAI & al. (1985)
Myrmecia sp. 2
2n = 68, Indonesia, IMAI & al. (1995)
Myrmica (Myrmicinae)
Myrmica laevinodis (see M. rubra)
Myrmica lobicornis
n = 24, HAUSCHTECK (1965), HAUSCHTECK-JUNGEN & JUNGEN (1983)
Myrmica rubida (see Manica rubida)
Myrmica rubra*
n = 23, 2n = 46, Japan, IMAI (1969)
2n = 48, Switzerland, HAUSCHTECK-JUNGEN & JUNGEN (1983, M)
Myrmica ruginodis
n = 24, 2n = 48, Switzerland, HAUSCHTECK (1965), HAUSCHTECK-JUNGEN & JUNGEN (1983, M)
Myrmica sabuleti
n = 23, 2n = 46, Switzerland, HAUSCHTECK (1965), HAUSCHTECK-JUNGEN & JUNGEN (1983)
Myrmica scabrinodis
n = 22, 2n = 44, Switzerland, HAUSCHTECK (1965)
Myrmica schencki
n = 23, 2n = 46, Switzerland, HAUSCHTECK-JUNGEN & JUNGEN (1983, M)
Myrmica sulcinodis*
n = 24-28 (HAUSCHTECK-JUNGEN, in CROZIER 1975)
n = 28, 2n = 56, Switzerland, HAUSCHTECK (1965, M), HAUSCHTECK-JUNGEN & JUNGEN (1983, M)
Myrmicaria (Myrmicinae)
Myrmicaria brunnea
2n = 44, India, IMAI & al. (1984, K)
Myrmicaria sp.
2n = 44, Indonesia, IMAI & al. (1985)
Myrmicaria sp. 1
2n = 44, Malaysia, GOСI & al. (1982)
Myrmicaria sp. 2
2n = 44, Malaysia, GOСI & al. (1982), IMAI & al. (1983)
Myrmicaria sp. 3
2n = 44, Malaysia, GOСI & al. (1982)
Myrmicaria sp. 4
n = 23, 2n = 46, Malaysia, IMAI & al. (1983)
Myrmoxenus (Myrmicinae)
Myrmoxenus adlerzi (Epimyrma adlerzi)
n = 10, Greece, DOUWES & al. (1988, K)
Myrmoxenus algeriana (Epimyrma algeriana)
n = 10, DOUWES & al. 1988)
Myrmoxenus bernardi (Epimyrma bernardi)
n = 10, Spain, BUSCHINGER & al. (1986), FISCHER (1987, K)
Myrmoxenus corsica (Epimyrma corsica)
n = 10, France, BUSCHINGER & al. (1986), FISCHER (1987)
Myrmoxenus foreli (see M. kraussei)
Myrmoxenus gordiagini (Epimyrma gordiagini)
n = 10, Croatia, BUSCHINGER & al. (1983, K), FISCHER (1987, K)
Myrmoxenus kraussei (Epimyrma kraussei, E. foreli, E. vandeli)
n = 10, France, Italy, Spain, BUSCHINGER & al. (1986, K), FISCHER (1987, K)
Myrmoxenus ravouxi (Epimyrma ravouxi)
n = 10, Italy, France, Switzerland, Germany, BUSCHINGER & al. (1986, K), FISCHER (1987, K)
Myrmoxenus stumperi (Epimyrma stumperi)
n = 10, France, Switzerland, BUSCHINGER & al. (1986), FISCHER (1987)
Myrmoxenus vandeli (see M. kraussei)
Mystrium (Amblyoponinae)
Mystrium camillae
2n = 32, Sarawak, TJAN & al. (1986)
Neivamyrmex (Ecitoninae)
Neivamyrmex texanus
2n = 36, USA, TABER & COKENDOLPHER (1988, K)
Nothomyrmecia (Myrmeciinae)
Nothomyrmecia macrops
2n = 94, Australia, IMAI & al. (1990, K)
Notoncus (Formicinae)
Notoncus ectatommoides
n = 22, 2n = 44, Australia, IMAI & al. (1977, K)
Ochetellus (Dolichoderinae)
Ochetellus glaber (O. itoi, Iridomyrmex glaber, I. itoi)
n = 14, 2n = 28, Australia, CROZIER (1968a, K)
n = 14, 2n = 28, Japan, IMAI & YOSIDA (1964), IMAI (1969, K)
Odontomachus (Ponerinae)
Odontomachus latidens*
n = 15, Malaysia, IMAI & al. (1983)
2n = 32, Indonesia, IMAI & al. (1985)
Odontomachus rixosus*
n = 15, 2n = 30, Malaysia, GOСI & al. (1982, as O. sp. 2), IMAI & al. (1983)
Odontomachus simillimus
n = 22, 2n = 44, Malaysia, GOСI & al. (1982, as O. sp. 1), IMAI & al. (1983)
n = 22, 2n = 44, Indonesia, IMAI & al. (1985)
2n = 44, Sarawak, TJAN & al. (1986)
Odontomachus sp. (ANIC-1)
2n = 44, Australia, IMAI & al. (1977, K)
Odontomachus sp. 3
n = 22, 2n = 44, Malaysia, GOСI & al. (1982)
Odontoponera (Ponerinae)
Odontoponera transversa*
2n = 46, India, IMAI & al. (1984, K)
2n = 42, Indonesia, IMAI & al. (1985)
Oecophylla (Formicinae)
Oecophylla longinoda
n = 12, LEDOUX (1954, in CROZIER 1970b)
Oecophylla smaragdina
n = 8, 2n = 16, Malaysia, CROZIER (1970b, K)
2n = 16, India, IMAI & al. (1984, K)
Oligomyrmex (see under Carebara)
Opisthopsis (Formicinae)
Opisthopsis rufithorax
2n = 50, Australia, IMAI & al. (1977, K)
Orectognathus (Myrmicinae)
Orectognathus clarki
2n = 30, Australia, CROZIER (1968d, M, K)
Orectognathus darlingtoni
n = 11, 2n = 22, Australia, IMAI & al. (1977, K)
Orectognathus versicolor
n = 11, 2n = 22, Australia, IMAI & al. (1977, K)
Pachycondyla (Ponerinae)
Pachycondyla arhuaca
2n = 12, Brazil, MARIANO & al. (2007)
Pachycondyla apicalis*
2n = 36, 40, 68, Brazil, MARIANO & al. (2007), DELABIE & al. (2008, M, K)
Pachycondyla astuta*
2n = 18, 22, Indonesia, IMAI & al. (1985)
Pachycondyla carinulata
2n = 24, Brazil, MARIANO & al. (2007)
Pachycondyla chinensis (Brachyponera chinensis)
2n = 22, Japan, IMAI & KUBOTA (1972, K)
Pachycondyla crassinoda
2n = 62, Brazil, MARIANO & al. (2006c, M), MARIANO & al. (2007)
Pachycondyla crenata
2n = 26, Brazil, MARIANO & al. (2006a, M), MARIANO & al. (2007)
Pachycondyla constricta
2n = 30, Brazil, MARIANO & al. (2007)
Pachycondyla gilberti
2n = 14, Brazil, MARIANO & al. (2007)
Pachycondyla goeldii
2n = 24, French Guyana, MARIANO & al. (2007)
Pachycondyla harpax
2n = 96, Brazil, MARIANO & al. (2006c, M), MARI-ANO & al. (2007)
Pachycondyla inversa
2n = 30, Brazil, MARIANO & al. (1999), MARIANO & al. (2007)
Pachycondyla leeuwenhoeki
n = 8, 2n = 16, Malaysia, IMAI & al. (1983)
Pachycondyla lutea (Brachyponera lutea)
2n = 16, Australia, IMAI & al. (1977, K)
Pachycondyla luteipes (Brachyponera luteipes)
n = 11, 2n = 22, Japan, IMAI & YOSIDA (1964), IMAI (1969, K)
2n = 22, India, IMAI & al. (1984, K)
Pachycondyla marginata
2n = 46, Brazil, MARIANO & al. (2007)
Pachycondyla mesonotalis
n = 13, 2n = 26, Brazil, MARIANO & al. (2006a, M), MARIANO & al. (2007)
Pachycondyla rubiginosa (Bothroponera rubiginosa)
2n = 76, India, IMAI & al. (1984, K)
Pachycondyla rubra*
2n = 38, 40, Malaysia, IMAI & al. (1983)
n = 10, 2n = 20, Indonesia, IMAI & al. (1985)
2n = 20, Sarawak, TJAN & al. (1986)
Pachycondyla rufipes (Bothroponera rufipes)
2n = 48, India, IMAI & al. (1984, K)
Pachycondyla sinensis (see P. chinensis)
Pachycondyla stigma
2n = 12, Brazil, MARIANO & al. (2007)
Pachycondyla striata
2n = 104, Brazil, MARIANO & al. (2007)
Pachycondyla "subversa"
2n = 26, 28, Brazil, MARIANO & al. (2007)
Pachycondyla tridentata
2n = 28, Malaysia, IMAI & al. (1983)
Pachycondyla unidentata
2n = 12, Brazil, MARIANO & al. (2007)
Pachycondyla venusta
2n = 48, Brazil, MARIANO & al. (2007)
Pachycondyla verenae*
2n = 42, 58-60, 62, 64, Brazil, MARIANO & al. (2007), DELABIE & al. (2008, M, K)
Pachycondyla villosa
2n = 34, Brazil, MARIANO & al. (1999), MARIANO & al. (2007)
Pachycondyla sp.
2n = 22, Malaysia, GOСI & al. (1982, as Brachyponera sp.)
Pachycondyla sp.
2n = 38, India, IMAI & al. (1984, K, as Ectomomyrmex sp.)
Pachycondyla sp. (near P. obscurans)
2n = 44, Indonesia, IMAI & al. (1985)
Pachycondyla sp.
2n = 60, Australia, IMAI & al. (1977, K, as Bothro-ponera sp. (ANIC-2)
Pachycondyla sp.
n = 11, 2n = 22, Malaysia, GOСI & al. (1982)
Pachycondyla sp. 1
2n = 28, Malaysia, GOСI & al. (1982, as Mesoponera sp. 1)
Pachycondyla sp. 2
2n = 22, Malaysia, GOСI & al. (1982, as Mesoponera sp. 2)
Pachycondyla sp. 3
2n = 36, Malaysia, GOСI & al. (1982, as Mesoponera sp. 3)
Pachycondyla sp.
n = 18, 2n = 36, Malaysia, IMAI & al. (1983)
Pachycondyla sp. 1
2n = 48, India, IMAI & al. (1984, K, as Bothroponera sp. 1, near tesserinoda)
Pachycondyla sp. 2
2n = 52, India, IMAI & al. (1984, K, as Bothroponera sp. 2, near P. tesserinoda)
Pachycondyla sp. (P. venusta gr.)
2n = 54, Brazil, MARIANO & al. (2007)
Papyrius (Dolichoderinae)
Papyrius nitidus (Iridomyrmex nitidus)
n = 8, 2n = 16, Australia, CROZIER (1968a, K), IMAI & al. (1977, K)
Paratrechina (Formicinae)
Paratrechina indica
2n = 30, India, IMAI & al. (1984, K )
Paratrechina longicornis*
2n = 16, Taiwan, HUNG & al. (1972)
n = 8, 2n = 16, India, IMAI & al. (1984, K)
n = 16, Indonesia, IMAI & al. (1985)
Paratrechina parvula
n = 15, HAUSCHTECK-JUNGEN (in CROZIER 1975)
Paratrechina sp. (ANIC-1)
2n = 30, Australia, IMAI & al. (1977, K)
Paratrechina sp. 1
2n = 16, Malaysia, GOСI & al. (1982)
Paratrechina sp. 2
2n = 26, Malaysia, GOСI & al. (1982)
Paratrechina sp. 3
2n = 28, Malaysia, GOСI & al. (1982)
Paratrechina sp. 4
n = 8, 2n = 16 Malaysia, GOСI & al. (1982), IMAI & al. (1983)
Paratrechina sp. 5
2n = 28, Malaysia, GOСI & al. (1982)
Paratrechina sp. 6
2n = 30, Malaysia, IMAI & al. (1983)
Paratrechina sp. 7
2n = 16, Malaysia, IMAI & al. (1983)
Paratrechina sp. 3 (near P. yerbuyi)
n = 15, 2n = 30, India, IMAI & al. (1984, K)
Paratrechina sp. 2
2n = 30, Indonesia, IMAI & al. (1985)
Paratrechina sp. 3
2n = 30, Indonesia, IMAI & al. (1985)
Pheidole (Myrmicinae)
Pheidole binghamii
2n = 20, Indonesia, IMAI & al. (1985)
Pheidole capellinii
2n = 20, Indonesia, IMAI & al. (1985)
Pheidole cornutula (see P. subarmata)
Pheidole dentata
2n = 20, USA, CROZIER (1970b, K)
Pheidole dentigula
2n = 20, USA, CROZIER (1970b, K)
Pheidole desertorum
2n = 20, USA, TABER & COKENDOLPHER (1988, K)
Pheidole fallax
n = 10, 2n = 20, Uruguay, GOСI & al. (1983, K)
Pheidole fervida
n = 10, 2n = 20, Japan, IMAI & YOSIDA (1964), IMAI (1966a, M), IMAI (1969)
Pheidole hortensis
2n = 20, Indonesia, IMAI & al. (1985)
Pheidole hyatti
2n = 20, USA, TABER & COKENDOLPHER (1988, K)
Pheidole indica (P. rotschana)
2n = 20, India, IMAI & al. (1984, K, as P. indica)
2n = 20, India, IMAI & al. (1984, K, as P. rotschana)
Pheidole latinoda
2n = 42, India, IMAI & al. (1984, K)
Pheidole megacephala
2n = 20, Malaysia, IMAI & al. (1983)
Pheidole mus
2n = 12, India, IMAI & al. (1984, K)
Pheidole nitidula (P. strobeli)
2n = 20, Uruguay, GOСI & al. (1983, K)
Pheidole noda*
n = 17-20, CROZIER (1975, K)
n = 19, 2n = 38, Japan, IMAI (1969)
n = 17-20, 2n = 37-39, Japan, IMAI & KUBOTA (1972, K)
Pheidole pallidula*
2n = 24, Switzerland, HAUSCHTECK (1961, M)
n = 10, 2n = 20, Switzerland, Spain, Greece, Tunisia, HAUSCHTECK-JUNGEN & JUNGEN (1983, K)
n = 10, 2n = 20, Spain, PALOMEQUE & al. (1987, M, K), PALOMEQUE & al. (1990b, M, K), LORITE & al. (2000, M. K)
Pheidole plagiaria
2n = 20, Indonesia, IMAI & al. (1985)
Pheidole porcula
2n = 20, USA, TABER & COKENDOLPHER (1988, K)
Pheidole rotschana (see P. indica)
Pheidole sitarches (see P. soritis)
Pheidole soritis (P. sitarches)
2n = 18, USA, TABER & COKENDOLPHER (1988, K)
Pheidole spininodis
2n = 20, Uruguay, GOСI & al. (1983, K)
Pheidole strobeli (see P. nitidula)
Pheidole subarmata (P. cornutula)
2n = 20, Uruguay, GOСI & al. (1983, K)
Pheidole tepicana
2n = 18, USA, TABER & COKENDOLPHER (1988)
Pheidole woodmasoni
2n = 18, India, IMAI & al. (1984, K)
Pheidole sp. (P. concentrica group)
n = 9, Australia, CROZIER (1966)
Pheidole sp.
n = 10, Japan, IMAI (1969)
Pheidole sp. (ANIC-20)
2n = 20, Australia, IMAI & al. (1977, K)
Pheidole sp. (ANIC-21)
2n = 20, Australia, IMAI & al. (1977, K)
Pheidole sp. (ANIC-22)
2n = 20, Australia, IMAI & al. (1977, K)
Pheidole sp. (ANIC-23)
2n = 20, Australia, IMAI & al. (1977, K)
Pheidole sp. (ANIC-24)
2n = 18, Australia, IMAI & al. (1977, K)
Pheidole sp. (ANIC-25)
2n = 20, Australia, IMAI & al. (1977, K)
Pheidole sp. (ANIC-26)
2n = 20, Australia, IMAI & al. (1977, K)
Pheidole sp. (ANIC-27)
2n = 20, Australia, IMAI & al. (1977, K)
Pheidole sp. (ANIC-28)
2n = 20, Australia, IMAI & al. (1977, K)
Pheidole sp. (ANIC-29)
2n = 20, Australia, IMAI & al. (1977, K)
Pheidole sp. (ANIC-30)
2n = 20, Australia, IMAI & al. (1977, K)
Pheidole sp. (ANIC-31)
2n = 20, Australia, IMAI & al. (1977, K)
Pheidole sp. (ANIC-32)
2n = 20, Australia, IMAI & al. (1977, K)
Pheidole sp. 1
n = 10, 2n = 20, Malaysia, GOСI & al. (1982), IMAI & al. (1983)
Pheidole sp. 2
n = 10, 2n = 20, Malaysia, GOСI & al. (1982)
Pheidole sp. 3
2n = 20, Malaysia, GOСI & al. (1982)
Pheidole sp. 4
2n = 20, Malaysia, GOСI & al. (1982)
Pheidole sp. 5
n = 16-17, Malaysia, GOСI & al. (1982)
Pheidole sp. 6
2n = 20, Malaysia, GOСI & al. (1982)
Pheidole sp. 7
2n = 16, Malaysia, GOСI & al. (1982), IMAI & al. (1983)
Pheidole sp. 8
2n = 38, Malaysia, GOСI & al. (1982)
Pheidole sp. 9
2n = 18, Malaysia, GOСI & al. (1982)
Pheidole sp. 10
2n = 20, Malaysia, GOСI & al. (1982)
Pheidole sp. 11
2n = 22, Malaysia, IMAI & al. (1983)
Pheidole sp. 12
n = 10, 2n = 20, Malaysia, IMAI & al. (1983)
Pheidole sp. 13
2n = 38, Malaysia, IMAI & al. (1983)
Pheidole sp. 14
2n = 20, Malaysia, IMAI & al. (1983)
Pheidole sp. 15
2n = 20, Malaysia, IMAI & al. (1983)
Pheidole sp. 16
n = 9, 2n = 18, Malaysia, IMAI & al. (1983)
Pheidole sp. 17
2n = 20-21, Malaysia, IMAI & al. (1983)
Pheidole sp. 18
2n = 20, Malaysia, IMAI & al. (1983)
Pheidole sp. 19
2n = 20, Malaysia, IMAI & al. (1983)
Pheidole sp. 20
n = 18, 2n = 36, Malaysia, IMAI & al. (1983)
Pheidole sp. 1 (near P. grayi)
2n = 42, India, IMAI & al. (1984, K)
Pheidole sp. 3
2n = 20, India, IMAI & al. (1984, K)
Pheidole sp. 4
2n = 20, India, IMAI & al. (1984)
Pheidole sp. 5
2n = 30, India, IMAI & al. (1984, K)
Pheidole sp. 8
2n = 20, India, IMAI & al. (1984, K)
Pheidole sp. 9 (near P. fossulata)
2n = 20, India, IMAI & al. (1984, K)
Pheidole sp. 11
2n = 20, India, IMAI & al. (1984)
Pheidole sp. 13 (near P. watsoni)
2n = 28, India, IMAI & al. (1984, K)
Pheidole sp. 14
2n = 18, India, IMAI & al. (1984, K)
Pheidole sp. 5
2n = 18, Indonesia, IMAI & al. (1985)
Pheidole sp. 6
2n = 18, Indonesia, IMAI & al. (1985)
Pheidole sp. 7
2n = 18, Indonesia, IMAI & al. (1985)
Pheidole sp. 8
2n = 32, Indonesia, IMAI & al. (1985)
Pheidole sp. 1
n = 10, 2n = 20, Sarawak, TJAN & al. (1986)
Pheidole sp. 2
2n = 18, Sarawak, TJAN & al. (1986)
Pheidole sp. 3
n = 9, 2n = 18, Sarawak, TJAN & al. (1986)
Pheidole sp. 4
2n = 18, Sarawak, TJAN & al. (1986)
Pheidologeton (Myrmicinae)
Pheidologeton diversus
2n = 42, India, IMAI & al. (1984, K)
Pheidologeton sp.
2n = 42, Sarawak, TJAN & al. (1986)
Philidris (Dolichoderinae)
Philidris cordata (Iridomyrmex cordatus)
2n = 16, Malaysia, IMAI & al. (1983)
2n = 16, Sarawak, TJAN & al. (1986)
Plagiolepis (Formicinae)
Plagiolepis barbara (see P. schmitzii)
Plagiolepis pygmaea*
n = 9, 2n = 18, Spain, Switzerland, Croatia, HAUSCHTECK-JUNGEN & JUNGEN (1983, K)
n = 9, 2n = 18, Spain, PALOMEQUE & al. (1993a, M, K)
Plagiolepis schmitzii (P. barbara)*
n = 9, 2n = 18, Tunisia, HAUSCHTECK-JUNGEN & JUNGEN (1983, K)
n = 9, 2n = 18, Spain, PALOMEQUE & al. (1990b, M, K), PALOMEQUE & al. (1993a, M, K)
Plagiolepis sp.
2n = 18, Indonesia, IMAI & al. (1985)
Plagiolepis sp.
2n = 18, Malaysia, IMAI & al. (1983)
Platythyrea (Ponerinae)
Platythyrea quadridenta
n = 9, 2n = 18, Malaysia, IMAI & al. (1983)
Platythyrea tricuspidata*
2n = 92-94, Malaysia, IMAI & al. (1983)
Podomyrma (Myrmicinae)
Podomyrma adelaidae*
2n = 49-51, Australia, IMAI & al. (1977, K)
Pogonomyrmex (Myrmicinae)
Pogonomyrmex apache
2n = 32, USA, TABER & al. (1988, K)
Pogonomyrmex badius
2n = 32, USA, TABER & al. (1988, K)
Pogonomyrmex barbatus
n = 16, 2n = 32, USA, TABER & al. (1988, K)
Pogonomyrmex brevispinosus
2n = 32, USA, TABER & al. (1988, K)
Pogonomyrmex californicus (P. estebanius)
2n = 32, USA, TABER & al. (1988, K)
Pogonomyrmex comanche
n = 16, 2n = 32, USA, TABER & al. (1988, K)
Pogonomyrmex desertorum
2n = 32, USA, TABER & al. (1988, K)
Pogonomyrmex estebanius (see P. californicus)
Pogonomyrmex huachucanus
2n = 36, USA, TABER & al. (1988, K)
Pogonomyrmex imberbiculus*
n = 30, 2n = 58-62, USA, TABER & al. (1988, K)
Pogonomyrmex magnacanthus
2n = 32, USA, TABER & al. (1988, K)
Pogonomyrmex maricopa
n = 16, 2n = 32, USA, TABER & al. (1988, K)
Pogonomyrmex montanus
2n = 32, USA, TABER & al. (1988, K)
Pogonomyrmex occidentalis
2n = 32, USA, MEHLHOP & GARDNER (1982), TABER & al. (1988, K)
Pogonomyrmex rugosus
2n = 32, USA, TABER & al. (1988, K)
Pogonomyrmex subnitidus
n = 16, 2n = 32, USA, TABER & al. (1988, K)
Polyergus (Formicinae)
Polyergus samurai
n = 27, 2n = 54, Japan, IMAI & YOSIDA (1964), IMAI (1966, M), IMAI (1969)
Polyrhachis (Formicinae)
Polyrhachis ammon
n = 21, 2n = 42, Australia, IMAI & al. (1977, K)
Polyrhachis dives
n = 21, Taiwan, HUNG & al. (1972)
Polyrhachis gribodoi
2n = 48, Indonesia, IMAI & al. (1985)
Polyrhachis hector
n = 21, 2n = 42, Malaysia, IMAI & al. (1983)
Polyrhachis hippomanes
n = 20, 2n = 40, Japan, IMAI (1969, K)
Polyrhachis illaudata*
n = 18, Malaysia, IMAI & al. (1983)
n = 14, 2n = 28, Indonesia, IMAI & al. (1985)
Polyrhachis lacteipennis (P. simplex)
2n = 42, India, IMAI & al. (1984, K)
Polyrhachis lamellidens
2n = 42, Japan, IMAI (1969)
Polyrhachis rastellata
n = 21, 2n = 42 Malaysia, CROZIER (1970b, K), IMAI & al. (1983)
Polyrhachis simplex (see P. lacteipennis)
Polyrhachis sp. (ANIC-1)
2n = 42, Australia, IMAI & al. (1977, K)
Polyrhachis sp. 1
n = 21, Malaysia, GOСI & al. (1982)
Polyrhachis sp. 2
n = 21, Malaysia, GOСI & al. (1982)
Polyrhachis sp. 3
2n = 20, Malaysia, GOСI & al. (1982)
Ponera (Ponerinae)
Ponera japonica
2n = 12, Malaysia, IMAI & al. (1983)
Ponera pennsylvanica
2n = 12, USA, HAUSCHTECK-JUNGEN & JUNGEN (1983, M)
Ponera ruginoda (see Myrmecia pilosula)
Ponera scabra*
n = 3-4, 2n = 7-8, Japan, IMAI & KUBOTA (1972, K), CROZIER (1975, M), IMAI & al. (1988a, K)
Ponera sp.
n = 6, Japan, IMAI (1969, K)
Ponera sp.
2n = 12, Indonesia, IMAI & al. (1985)
Prenolepis (Formicinae)
Prenolepis imparis
2n = 16, Switzerland, HAUSCHTECK (1962, M)
Prenolepis jerdoni*
n = 16, 20, 25, 27, 2n = 30-32, 34, 36, Malaysia, GOСI & al. (1982), IMAI & al. (1983)
Pristomyrmex (Myrmicinae)
Pristomyrmex pungens (see P. punctatus)
Pristomyrmex punctatus (P. pungens)
n = 12, 2n = 24, Japan, IMAI & YOSIDA (1964), IMAI (1966, M), IMAI (1969), ITOW & al. (1984, M)
Pristomyrmex sp.
2n = 22, Malaysia, GOСI & al. (1982)
Pristomyrmex sp. 2
n = 14, Malaysia, IMAI & al. (1983)
Proatta (Myrmicinae)
Proatta sp.
2n = 32, Malaysia, GOСI & al. (1982)
Probolomyrmex (Proceratiinae)
Probolomyrmex sp.
2n = 28, Malaysia, GOСI & al. (1982)
Proceratium (Proceratiinae)
Proceratium silaceum
n = 18, 2n = 46, USA, CROZIER (1970b, K)
Proceratium sp.
2n = 48, Sarawak, TJAN & al. (1986)
Prolasius (Formicinae)
Prolasius sp. (ANIC-1)
n = 9, 2n = 18, Australia, IMAI & al. (1977, K)
Prolasius sp. (ANIC-2)
2n = 18, Australia, IMAI & al. (1977, K)
Protomognathus (Myrmicinae)
Protomognathus americanus (Harpagoxenus americanus)
n = 11, USA, Canada, FISCHER (1987, K)
Pseudolasius (Formicinae)
Pseudolasius sp. (near P. emeryi)
n = 14, Taiwan, HUNG & al. (1972)
Pseudolasius sp.
2n = 30, Indonesia, IMAI & al. (1985)
Pseudolasius sp.
2n = 30, Sarawak, TJAN & al. (1986)
Pseudolasius sp. 1
n = 8, Malaysia, IMAI & al. (1983)
Pseudolasius sp. 2*
n = 15, 17, 19, 2n = 30, Malaysia, IMAI & al. (1983)
Pseudomyrmex (Pseudomyrmecinae)
Pseudomyrmex gracilis
2n = 70, Brazil, SPOSITO & al. (2006)
Pseudomyrmex holmgreni
2n = 50, Brazil, SPOSITO & al. (2006)
Pseudomyrmex penetrator
2n = 24, Brazil, SPOSITO & al. (2006)
Pseudomyrmex schuppi
n = 24, Brazil, SPOSITO & al. (2006)
Pseudomyrmex sp. 1 nr. simplex
2n = 43, Brazil, SPOSITO & al. (2006)
Pseudomyrmex sp. 2 nr. simplex
2n = 50, Brazil, SPOSITO & al. (2006)
Pseudomyrmex sp. 3
2n = 44, Brazil, SPOSITO & al. (2006)
Pyramica (Myrmicinae)
Pyramica dohertyi (Smithistruma taipingensis)
n = 12, 2n = 24, Malaysia, IMAI & al. (1983)
Pyramica mutica (Kyidris mutica)
2n = 36, Indonesia, IMAI & al. (1985)
Pyramica sp.
2n = 38, Indonesia, IMAI & al. (1985, as Smithistruma sp.)
Pyramica sp.
n = 13, BROWN (In CROZIER 1975, as Smithistruma sp.)
Pyramica sp.
2n = 16, Malaysia, GOСI & al. (1982, as Smithistruma sp.)
Recurvidris (Myrmicinae)
Recurvidris sp. (Trigonogaster sp.)
2n = 24, India, IMAI & al. (1984, K)
Rhytidoponera (Ectatomminae)
Rhytidoponera aciculata
2n = 52, Australia, IMAI & al. (1977, K)
Rhytidoponera chalybaea
2n = 42, Australia, IMAI & al. (1977, K)
Rhytidoponera impressa
2n = 42, Australia, IMAI & al. (1977, K)
Rhytidoponera lamellinodis
2n = 52, Australia, CROZIER & al. (1986)
Rhytidoponera maniae*
2n = 39, 44, 48, Australia, IMAI & al. (1977, K)
Rhytidoponera mayri
2n = 50, Australia, IMAI & al. (1977, K)
Rhytidoponera metallica*
n = 17-22, 2n = 22-24, 36-37, 41-44, 46, Australia, IMAI & al. (1977)
2n = 24, 41-43, Australia, CROZIER (1969, K, M)
2n = 24, 34-38, 46, Australia, CROZIER & al. (1986)
Rhytidoponera punctata
2n = 50, Australia, CROZIER & al. (1986)
Rhytidoponera purpurea
2n = 38, Australia, IMAI & al. (1977, K)
Rhytidoponera tasmaniensis-1
2n = 30, Australia, CROZIER & al. (1986)
Rhytidoponera tasmaniensis-2
2n = 46, Australia, CROZIER & al. (1986)
Rhytidoponera victoriae
n = 21, 2n = 42, Australia, CROZIER (1969, M), CROZIER (1970b), CROZIER & al. (1986), IMAI & al. (1977, K)
Rhytidoponera sp. (ANIC-9)
2n = 48, Australia, CROZIER & al. (1986)
Rhytidoponera sp. (ANIC-10)
2n = 46, 49, Australia, CROZIER & al. (1986)
Rhytidoponera sp. (ANIC-11)
2n = 50, Australia, CROZIER & al. (1986)
Rhytidoponera sp. (ANIC-12)
2n = 50, Australia, CROZIER & al. (1986)
Rhytidoponera sp. (ANIC-13)
2n = 52, Australia, CROZIER & al. (1986)
Rhytidoponera sp. (ANIC-14)
2n = 23, Australia, CROZIER & al. (1986)
Rhytidoponera sp. (ANIC-15)
2n = 50, Australia, CROZIER & al. (1986)
Rhytidoponera sp. (ANIC-16)
2n = 52, Australia, CROZIER & al. (1986)
Sericomyrmex (Myrmicinae)
Sericomyrmex amabilis
2n = 50, Panama, MURAKAMI & al. (1998, K)
Smithistruma (see under Pyramica)
Solenopsis (Myrmicinae)
Solenopsis aurea
2n = 32, USA, CROZIER (1970b, K)
Solenopsis fugax
n = 11, 2n = 22, Switzerland, HAUSCHTECK (1961, M)
2n = 22, Japan, IMAI (1969, K)
Solenopsis geminata
n = 16, 2n = 32, USA, CROZIER (1970b, K)
2n = 32, India, IMAI & al. (1984, K)
Solenopsis invicta
n = 16, 2n = 32, USA, GLANCEY & al. (1976)
Solenopsis molesta
2n = 22, USA, CROZIER (1970b, K)
Solenopsis richteri
n = 16, USA, GLANCEY & al. (1976)
Solenopsis saevissima
n = 16, 2n = 32, Uruguay, GOСI & al. (1983, K)
Solenopsis xyloni
n = 16, 2n = 32, USA, TABER & COKENDOLPHER (1988, K)
Solenopsis sp.
2n = 38, Malaysia, GOСI & al. (1982, as Diplrhoptrum sp.)
Solenopsis sp.
n = 11, Australia, CROZIER (1966)
Sphinctomyrmex (Cerapachyinae)
Sphinctomyrmex steinheili*
2n = 45-46, Australia, IMAI & al. (1977, K)
Stenamma (Myrmicinae)
Stenamma brevicorne
n = 4, 2n = 8, USA, HAUSCHTECK (1962, M), HAUSCHTECK (1963)
Stenamma westwoodii
n = 20, HAUSCHTECK-JUNGEN (In CROZIER 1975)
Stigmacros (Formicinae)
Stigmacros sp. (ANIC-1)
2n = 38, Australia, IMAI & al. (1977, K)
Stigmacros sp. (ANIC-3)
n = 10, 2n = 20, Australia, IMAI & al. (1977, K)
Strongylognathus (Myrmicinae)
Strongylognathus huberi
n = 14, Switzerland, HAUSCHTECK (1962, M)
Strumigenys (Myrmicinae)
Strumigenys doriae
2n = 22, Indonesia, IMAI & al. (1985)
Strumigenys friedae
2n = 24, Australia, IMAI & al. (1977, K)
Strumigenys godeffroyi*
2n = 40, Malaysia, IMAI & al. (1983)
2n = 44, Indonesia, IMAI & al. (1985)
Symmyrmica (see under Formicoxenus)
Tapinoma (Dolichoderinae)
Tapinoma erraticum*
n = 8, 2n = 16, Switzerland, HAUSCHTECK-JUNGEN & JUNGEN (1983, K)
n = 8, 2n = 16, Spain, PALOMEQUE & al. (1988, M, K), LORITE & al. (1998b, M, K)
Tapinoma indicum*
n = 7, IMAI (in CROZIER 1975)
2n = 10, India, IMAI & al. (1984, K)
Tapinoma melanocephalum
n = 5, 2n = 10, USA, CROZIER (1970a, K) (1970b)
2n = 10, India, IMAI & al. (1984, K)
2n = 10, Indonesia, IMAI & al. (1985)
Tapinoma nigerrimum*
n = 9, Tunisia, HAUSCHTECK-JUNGEN & JUNGEN (1983, M)
n = 9, 2n = 18 (Spain), PALOMEQUE & al. (1988, M, K), PALOMEQUE & al. (1990a, M), PALOMEQUE & al. (1990b, M, K), PALOMEQUE & al. (1993a, M, K), LORITE & al. (1996a, M, K), LORITE & al. (1997, M) ), LORITE & al. (1998b, M, K), LORITE & al. (1999a, M, K), LORITE & al. (1999b, M, K), LORITE & al. (2002a, M, K)
Tapinoma sessile
n = 8, 2n = 16, USA, CROZIER (1970a, M, K), TABER & COKENDOLPHER (1988, K)
Tapinoma simrothi
n = 9, HAUSCHTECK-JUNGEN (in CROZIER, 1975)
2n = 18, Tunisia, Spain, HAUSCHTECK-JUNGEN & JUNGEN (1983)
Tapinoma sp.
n = 5, 2n = 10, Malaysia, GOСI & al. (1982)
Technomyrmex (Dolichoderinae)
Technomyrmex albipes*
2n = 16, Australia, IMAI & al. (1977, K)
2n = 16, India, IMAI & al. (1984, K)
n = 9, 2n = 18, Australia, CROZIER (1968a, K)
Technomyrmex sp. 2 (T. bicolor group)
2n = 28, India, IMAI & al. (1984, K)
Technomyrmex sp.
2n = 30, Malaysia, GOСI & al. (1982), IMAI & al. (1983)
Technomyrmex sp. 1
2n = 28, Indonesia, IMAI & al. (1985)
Technomyrmex sp. 2
2n = 30, Indonesia, IMAI & al. (1985)
Temnothorax (Myrmicinae)
Temnothorax affinis (Leptothorax affinis)
n = 9, France, Germany, Switzerland, FISCHER (1987, K)
Temnothorax albipennis (Leptothorax albipennis, L. tuberointerruptus, Stenamma albipennis)*
n = 8, Great Britain, ORLEDGE (1998, M, K)
n = 8, Germany, FISCHER (1987, K)
Temnothorax ambiguus (Leptothorax ambiguus)
n = 22, Canada, FISCHER (1987, K)
Temnothorax andrei (Leptothorax andrei)
n = 10, USA, FISCHER (1987, K)
Temnothorax angustulus (Leptothorax angustulus)
n = 23, France, FISCHER (1987, K, M)
Temnothorax congruus (Leptothorax congruus)
n = 9, 2n = 18, Japan, IMAI & KUBOTA (1972, K)
Temnothorax corticalis (Leptothorax corticalis)*
n = 9, HAUSCHTECK-JUNGEN (in CROZIER 1975)
n = 12, Germany, FISCHER (1987, K)
Temnothorax curvispinosus (Leptothorax curvispinosus)*
n = 23, USA, FISCHER (1987, M)
Temnothorax duloticus (Leptothorax duloticus)
n = 23-25, Canada, FISCHER (1987)
Temnothorax exilis (Leptothorax exilis)
n = 11, France, Croatia, Italy, Turkey, FISCHER (1987, K)
Temnothorax flavicornis (Leptothorax flavicornis)
n = 11, Italy, FISCHER (1987, K)
Temnothorax gredosi (Leptothorax gredosi)
n = 17, Spain, FISCHER (1987, K)
Temnothorax interruptus (Leptothorax interruptus)
2n = 24, Germany, HAUSCHTECK-JUNGEN & JUNGEN (1983, K)
n = 12, Germany, Croatia, FISCHER (1987, K)
Temnothorax kraussei (Leptothorax kraussei)
n = 14, France, FISCHER (1987, K)
Temnothorax lichtensteini (Leptothorax lichtensteini)
n = 14, France, Italy, Croatia, FISCHER (1987, K)
Temnothorax longispinosus (Leptothorax longispinosus)
n = 12, 2n = 24, USA, CROZIER (1970b, K)
n = 12, Canada FISCHER (1987, K)
Temnothorax melas (Leptothorax melas)
n = 13, France, FISCHER (1987, K)
Temnothorax niger (Leptothorax niger)
n = 18, France, Italy, FISCHER (1987, K)
Temnothorax nigriceps (Leptothorax nigriceps)*
2n = 18, Switzerland, HAUSCHTECK (1962, M)
n = 9, Italy, Croatia, France, Germany, FISCHER (1987, K)
Temnothorax nylanderi (Leptothorax nylanderi)
n = 11, 2n = 22, Switzerland, HAUSCHTECK-JUNGEN & JUNGEN (1983)
n = 11, Germany, Croatia, France, FISCHER (1987, K)
Temnothorax parvulus (Leptothorax parvulus)
n = 14, (Italy), France, Switzerland, FISCHER (1987, K)
Temnothorax rabaudi (Leptothorax rabaudi)
n = 9, France, FISCHER (1987, K)
Temnothorax racovitzai (Leptothorax racovitzai)
n = 21, France, FISCHER (1987, K)
Temnothorax recedens (Leptothorax recedens)
n = 12, Italy, FISCHER (1987, K)
Temnothorax rottenbergii (Leptothorax rottenbergii)
n = 11, Italy, FISCHER (1987, K)
Temnothorax rougeti (see T. unifasciatus)
Temnothorax rugatulus (Leptothorax rugatulus)*
n = 14, 2n = 26-27, USA, TABER & COKENDOLPHER (1988, K), FISCHER (1987, K)
Temnothorax schaumii (Leptothorax schaumii)
n = 9, USA, HAUSCHTECK-JUNGEN & JUNGEN (1983)
Temnothorax sordidulus (Leptothorax sordidulus)
n = 11, Italy, Croatia, Switzerland, FISCHER (1987, K)
Temnothorax specularis (Leptothorax specularis)
n = 17, Italy, FISCHER (1987, K)
Temnothorax spinosior (Leptothorax spinosior)*
n = 12, 2n = 24, Japan, IMAI (1966, M), IMAI (1969, K), IMAI (1974, K), IMAI & YOSIDA (1964), FISCHER (1987, K)
Temnothorax spinosus (Leptothorax spinosus)
n = 16, Algeria, FISCHER (1987, K)
Temnothorax tristis (Leptothorax tristis)
n = 21, France, Switzerland, FISCHER (1987, K)
Temnothorax tuberum (Leptothorax tuberum)
n = 9, 2n = 18, Switzerland, HAUSCHTECK (1962, M, K), HAUSCHTECK-JUNGEN & JUNGEN (1983, K)
n = 9, Switzerland, Sweden, France, FISCHER (1987, K)
Temnothorax unifasciatus (Leptothorax rougeti, L. uni-fasciatus)*
n = 8, FISCHER (1987, K, as Leptothorax rougeti)
n = 9, 2n = 18, Germany, Switzerland, HAUSCHTECK-JUNGEN & JUNGEN (1983, K, as Leptothorax unifas-ciatus)
n = 9, Germany, France, Italy, Croatia, Greece, FISCHER (1987, K, as Leptothorax unifasciatus)
Temnothorax cf. lichtensteini (Leptothorax cf. lichtensteini)
n = 14, France, Spain, FISCHER (1987)
Temnothorax cf. interrupus (Leptothorax cf. interruptus)
n = 12, Greece, FISCHER (1987, K)
Temnothorax spp. (T. tuberum group)
n = 9, Greece, Croatia, France, FISCHER (1987, K, as Leptothorax spp., tuberum group)
Tetramorium (Myrmicinae)
Tetramorium adelphon
2n = 22, Indonesia, IMAI & al. (1985)
Tetramorium brevidentatum
n = 10, 2n = 20, Indonesia, IMAI & al. (1985)
Tetramorium caespitum
n = 14, 2n = 28, Switzerland, HAUSCHTECK (1961, M)
n = 14, 2n = 28, Japan, IMAI (1966, M), IMAI (1969, K), IMAI & YOSIDA (1964)
n = 14, Spain, PALOMEQUE & al. (1987, M, K), PALOMEQUE & al. (1990b, M, K), LORITE & al. (2000, M, K)
Tetramorium eleates
2n = 28, Malaysia, IMAI & al. (1983)
Tetramorium forte (T. hispanicum)*
n = 14, Spain, PALOMEQUE & al. (1987, M, K), LORITE & al. (2000, M, K)
Tetramorium guineense
2n = 22, Japan, IMAI (1969, K)
Tetramorium hispanicum (see Tetramorium forte)
Tetramorium insolens
2n = 22, Indonesia, IMAI & al. (1985)
Tetramorium kheperra
2n = 14, Indonesia, IMAI & al. (1985)
Tetramorium lanuginosum (Triglyphothrix lanuginosa)
2n = 14, India, IMAI & al. (1984, K)
Tetramorium pacificum
2n = 22, Indonesia, IMAI & al. (1985)
Tetramorium pnyxis
2n = 20, Malaysia, IMAI & al. (1983)
Tetramorium semilaeve
n = 14, Spain, PALOMEQUE & al. (1987, M, K), PALO-MEQUE & al. (1990b, M, K), LORITE & al. (2000, M, K)
Tetramorium seneb
2n = 20, Malaysia, IMAI & al. (1983)
Tetramorium simillimum
2n = 14, India, IMAI & al. (1984, K)
Tetramorium smithi
2n = 26, India, IMAI & al. (1984, K)
2n = 26, Indonesia, IMAI & al. (1985)
2n = 26, Sarawak, TJAN & al. (1986)
Tetramorium spinosum
n = 13, 2n = 26, USA, TABER & COKENDOLPHER (1988, K)
Tetramorium walshi
2n = 14, India, IMAI & al. (1984, K, as Triglyphothrix walshi)
Tetramorium sp.
2n = 20, Taiwan, HUNG & al. (1972, as Xiphomyrmex sp.)
Tetramorium sp. (ANIC-2)
2n = 18, Australia, IMAI & al. (1977, K, as Xiphomyr-mex sp. ANIC-2)
Tetramorium sp. ( ANIC-3)
2n = 20, Australia, IMAI & al. (1977, K, as Xiphomyr-mex sp. ANIC-3)
Tetramorium sp. (ANIC-4)
2n = 18, Australia, IMAI & al. (1977, K, as Xiphomyr-mex sp. ANIC-4)
Tetramorium sp. 1
2n = 20, Malaysia, GOСI & al. (1982, as Triglyphothrix sp. 1)
Tetramorium sp. 1
2n = 22, Malaysia, GOСI & al. (1982)
Tetramorium sp. 2
2n = 18, Malaysia, GOСI & al. (1982, as Triglyphothrix sp. 2)
Tetramorium sp. 3
2n = 26, Malaysia, GOСI & al. (1982)
Tetramorium sp. 4
2n = 14, Malaysia, GOСI & al. (1982)
Tetramorium sp. 3
2n = 35-36, India, IMAI & al. (1984, K)
Tetramorium sp. 2
2n = 20, Sarawak, TJAN & al. (1986)
Tetramorium sp. 3
2n = 24, Sarawak, TJAN & al. (1986)
Tetraponera (Pseudomyrmecinae)
Tetraponera allaborans
n = 16, Taiwan, HUNG & al. (1972)
Tetraponera sp.
2n = 44, Malaysia, GOСI & al. (1982)
Tetraponera sp. 2
2n = 42, Malaysia, IMAI & al. (1983)
Trachymyrmex (Myrmicinae)
Trachymyrmex septentrionales
n = 10, 2n = 20, Panama, MURAKAMI & al. (1998, K)
Trachymyrmex sp. 1
n = 6, 2n = 12, Panama, MURAKAMI & al. (1998, K)
Trachymyrmex sp. 2
2n = 18, Panama, MURAKAMI & al. (1998, K)
Triglyphothrix (see under Tetramorium)
Trigonogaster (see under Recurvidris)
Typhlomyrmex (Ectatomminae)
Typhlomyrmex meire
n = 10, 2n = 20, Brazil, MARIANO & al. (2006b, K)
Typhlomyrmex rogenhoferi
n = 17, 19, 2n = 34, 38, Brazil, MARIANO & al. (2006b, K)
2n = 36, French Guyana, MARIANO & al. (2006b, K)
Veromessor (see under Messor)
Vollenhovia (Myrmicinae)
Vollenhovia emeryii
n = 18, 2n = 36, Japan, IMAI & YOSIDA (1964), IMAI (1966a, M), IMAI (1969, K)
Vollenhovia sp.
n = 11, BROWN (in CROZIER 1975)
Vollenhovia sp. (ANIC-3)
2n = 40, Australia, IMAI & al. (1977, K)
Vollenhovia sp. 1
2n = 49-50, Malaysia, IMAI & al. (1983)
Vollenhovia sp. 2
2n = 33-34, Malaysia, IMAI & al. (1983)
Vollenhovia sp. 2
2n = 36, Indonesia, IMAI & al. (1985)
Xiphomyrmex (see under Tetramorium)
ПРИМЕЧАНИЯ ПО ВИДАМ (Appendix notes)
ПРИЛОЖЕНИЕ к ОСНОВНОЙ СТАТЬЕ.
Acropyga acutiventris: n = 15, 2n = 28-29 (GOСI & al. 1982, IMAI & al.1983). Only the chromosome number is reported in both papers, so that it is not possible to know the origin of the chromosome-number variability.
Anochetus graeffei: Two different chromosome numbers have been reported, 2n = 30 and 2n = 38 (IMAI & al. 1984, 1985). It is not possible to see the relationship since only the 2n = 30 karyotype has been published (IMAI & al. 1984).
Aphaenogaster beccarii: Two different chromosome num-bers have been reported for this species: 2n = 30 (IMAI & al. 1983) and 2n = 46 (IMAI & al. 1984). However, only the chromosome number was given in the first paper and the possible origin of this variation is not ex-plained.
Aphaenogaster gibbosa: For Swiss populations, n = 11 is reported (HAUSCHTECK-JUNGEN & JUNGEN 1983) but, in Spanish populations, n = 17 was found (PALOME-QUE & al. 1993b). It is not possible to make compari-sons since only the chromosome number was given in the first paper. In addition to this, in Spanish populations a polymorphism was also detected. Although the standard karyotype is n = 17, there are individuals with n = 16 due to a fusion of two telocentric chromosomes (LORITE & al. 2000).
Aphaenogaster iberica: In this species an intrapopulation polymorphism has been detected due to the presence of a supernumerary chromosome segment in the long arm of one of its metacentric chromosomes (PALOMEQUE & al. 1993a). The presence of this segment changes the chromosome to submetacentric.
Aphaenogaster longiceps: 2n = 45-46 (IMAI & al. 1977). In the 2n = 45 karyotype a heteromorphic chromosome pair is present; a large metacentric is single and there are two telocentric ones corresponding to its two arms.
Aphaenogaster rudis: Highly variable chromosome numbers have been found for this species, n = 16-18, 20,22 and 2n = 40-42, 44 (CROZIER 1969b, 1975, 1977). The different karyotypes are hardly related and it is possible that several sibling species are included in this taxon.
Aphaenogaster senilis: This species presents a polymor-phism similar to that observed in Aphaenogaster iberica (PALOMEQUE & al. 1993a) (see note).
Aphaenogaster smythiesi: Two different chromosome num-bers have been reported. In Japanese populations the diploid number is 2n = 22 with a karyotype formula n = 10 SM + 1 M (IMAI & al. 1971). However, Indian populations are 2n = 34, n = 8 M + 9 A (IMAI & al. 1984). The two karyotypes are difficult to relate but there are not enough data to determine whether they represent different species.
Aphaenogaster testaceopilosa: HAUSCHTECK-JUNGEN & JUNGEN (1983) indicated that material of this species was collected in Tunisia, Spain, and Croatia. Probably there are mistakes in some of the specimen identifications concerning the distribution of this species is North Africa (Alberto Tinaut, personal communication). The authors do not indicate what populations have been used to determine the chromosome number.
Camponotus alii: HAUSCHTECK-JUNGEN & JUNGEN (1983) reported the chromosome number of this species from Spanish populations. There is probably a mistake in the specimen identification since the presence of this species in Spain has not been cited (Alberto Tinaut, personal communication).
Camponotus compressus: Two different chromosome num-bers have been given for this species, n = 10 in Indian population (KUMBKARNI 1965) and n = 20 in Tunisian population (HAUSCHTECK-JUNGEN in CROZIER 1975). This was explained as a possible geographic variation. However, HAUSCHTECK-JUNGEN & JUNGEN (1983) indicated that the paper of KUMBKARNI (1965), has no convincing photographs and that their preparation show clearly n = 20 in the Tunisian specimens. Therefore, n = 10 is probably a mistake.
Camponotus crassisquamis: 2n = 39-40 (IMAI & al. 1984). The diploid number 2n = 39 is due to the presence of a heteromorphic chromosome pair with a metacentric and two telocentric chromosomes.
Camponotus cruentatus: HAUSCHTECK-JUNGEN & JUN-GEN (1983) reported n = 18 in a Spanish population whereas LORITE & al. (2002b) found n = 20. In the first paper, no photographs were provided, so that comparisons are not possible.
Camponotus japonicus: For this species two different chromosomes numbers have been published, n = 13 (IMAI & KUBOTA 1972) and n = 14 (IMAI 1966, 1969, IMAI & YOSIDA 1964). The analysis of the photograph in the paper of IMAI & KUBOTA (1972) shows that 13 chromosomes seem to be present. Thus, the value n = 14 could be a mistake due to the presence of a remarkable secondary constriction at the proximal part of the short arm of one chromosome. Probably this chromosome-satellite has been considered a small independent chromosome.
Camponotus rufipes: MARIANO & al. (2001) found in females a 2n = 39-40. The karyotype 2n = 39 presents a heteromorphic chromosome pair with a metacentric and two telocentric chromosomes.
Camponotus rufoglaucus: HAUSCHTECK-JUNGEN & JUN-GEN (1983) indicated that material of this species was collected in Tunisia and Spain. However, the presence of this species in Spain has not been cited (Alberto Tinaut, pers. comm.). The authors do not indicate whether the chromosome numbers have been derived from the analysis of the Tunisian or Spanish populations. Some Spanish myrmecologists consider that the Spanish cita-tions of this species belong to Camponotus micans (GУMEZ & ESPADALER 2009).
Camponotus (Myrmobrachys) sp., MARIANO & al. (2001): The diploid chromosome number of this species is 2n = 40, but the presence of one to three B-chromosomes has been detected.
Cataglyphis iberica: HAUSCHTECK-JUNGEN & JUNGEN (1983) reported the chromosome number of populations of Cataglypis albicans collected in Spain. However, because of the review of the genus Cataglyphis in Spain, the Spanish material classified as C. albicans may be reasigned to Cataglyphis iberica (TINAUT & PLAZA 1990).
Dinoponera lucida: The first chromosome number reported for this species was 2n = 106, making this the ant species with the highest chromosome number (MA-RIANO & al. 2004). With the analysis of new populations, higher chromosome numbers were found, 2n = 116, 118 and 120 (MARIANO & al. 2008), but the small size of the chromosomes do not allow the cytogenetic causes of the observed variation to be determined.
Dolichoderus thoracicus: 2n = 30-33 (IMAI & al. 1983, 1985). The variation in the chromosome number is due to the presence of B-chromosomes.
Formica truncorum: For this species an n = 26 was found in populations collected in Japan and in Switzerland (IMAI & YOSIDA 1964, IMAI 1969, HAUSCHTECK-JUNGEN & JUNGEN 1976). Later, in populations from Finland, ROSENGREN & al. (1980) found populations with n = 26 and also with n = 28, but the small size of the chromosomes of this species did not enable the or-igin of this variation to be determined.
Formicoxenus hirticolis: 2n = 30-34 (FRANCOEUR & al. 1985). Only the chromosome number was reported but the origin of the variation was not indicated.
Iridomyrmex anceps: Very different chromosome numbers have been published to this species; 2n = 18 in India (IMAI & al. 1984) and 2n = 48 in Indonesia (IMAI & al. 1985). Only the karyotype is available for the Indian specimens so that it is not possible to ascertain whether this variation is real or a mistake. In any case the 2n = 48 is not a usual chromosome number in the subfamily Dolichoderinae, which generally presents smaller chromosome numbers.
Iridomyrmex purpureus: In Australian populations CRO-ZIER (1968a, 1968c) found two different karyotypes. In the Beaumaris sample n = 6M + 1SM + 2A and in the Euston population n = 6M + 3SM. IMAI & al. (1977) analysed other Australian populations and found also the presence of these two karyotypes.
Iridomyrmex sp ANIC-5, CROZIER (1968a): This is the unique species under the genus Iridomyrmex with a chro-mosome number different from n = 9. The haploid chromosome number in this species is n = 7 (CROZIER 1968a). CROZIER (1968a) suggested that according to its karyotype this species could be removed from the genus Iridomyrmex. In fact with the review of the genus Iridomyrmex carried out of SHATTUCK (1992), all species with a chromosome number different of n = 9 were transferred to other genera.
Lasius alienus: Two different chromosome numbers have been found in this species, n = 14 (HAUSCHTECK 1962) and n = 15 (HAUSCHTECK-JUNGEN & JUNGEN 1983). This is presumably an instance of Robertsonian polymorphism, since in the n = 14 karyotype a sub-metacentric chromosome is present that is absent in the n = 15 karyotype.
Lasius brunneus: The standard chromosome number of this species is n = 15 (LORITE & al. 1998a) but a poly-morphism has been reported due to the presence of B-chromosomes (LORITE & al. 2002b).
Lasius niger: The standard chromosome number of this species is n = 15, 2n = 30. In a Spanish population a polymorphism due to the presence of B-chromosomes was detected (PALOMEQUE & al. 1990c).
Leptogenys diminuta: Two different chromosome num-bers have been reported in this species; 2n = 38 in Malaysian and Indian populations (GOСI & al. 1982, IMAI & al. 1983, 1984) and 2n = 32 in Indonesian populations (IMAI & al. 1985). The karyotype has been published only for the 2n = 38 karyotype (IMAI & al. 1984), so that it is not possible to determinate their relation with the 2n = 32 karyotype.
Leptothorax acervorum: In this species the standard karyotype is n = 13, although FISCHER (1987) found a polymorphism by a centric fission. These individuals have n = 14.
Leptothorax crassipilis: The variation n = 17 - 18 (FRANCOEUR 1986, LOISELLE & al. 1990) is probably due to Robertsonian polymorphism.
Leptothorax kutteri: The variation in the chromosome numbers (n = 23-25) found by BUSCHINGER & FISCHER (1991) has been explained as polymorphism by Robertsonian rearrangements.
Leptothorax muscorum: A high variation in the chromosome numbers was found in this species with n = 16 -23 (LOISELLE & al. 1990). In addition to this, some populations with the standard 17 - 18 chromosomes also present 4 to 7 B-chromosomes. LOISELLE & al. (1990) indicated that at least four species are under the taxon "muscorum".
Leptothorax retractus: The variation n = 17 - 18 (FRANCOEUR 1986, LOISELLE & al. 1990) was indicated as having been originated by Robertsonian rearrangements, although this is not clear. Nevertheless, there are some differences in the male genitalia of the two types of populations.
Monomorium indicum: The variation 2n = 21 - 22 (IMAI & al. 1984) is due to a reciprocal translocation polymorphism. The 2n = 22 seems to be the standard karyotype, since it is homomorphic and there are several heteromorphic karyotypes with 2n = 21. This polymor-phism has generated the presence of individuals with partial monosomy, trisomy and even tetrasomy.
Myrmecia banksi: See Myrmecia pilosula.
Myrmecia croslandi: See Myrmecia pilosula.
Myrmecia forficata: See Myrmecia pilosula.
Myrmecia fulvipes: See Myrmecia pilosula.
Myrmecia haskinsorum: See Myrmecia pilosula.
Myrmecia imaii: See Myrmecia pilosula.
Myrmecia mandibularis: See Myrmecia pilosula.
Myrmecia piliventris: See Myrmecia pilosula.
Myrmecia pilosula complex/ M. pilosula (s. str.): In this taxon, as in other Myrmecia species, the chromosome number frequently varies. Generally these variations are due to Robertsonian polymorphisms of centric fusion-fission, although other types of polymorphisms have been detected, such as pericentromeric inversions, translocations, deletions or heterochromatin growth (IMAI & al. 1988). In the taxon Myrmecia pilosula the haploid chromosome number found ranges between n = 1 to 2n = 32 (CROSLAND & CROZIER 1986, IMAI & al. 1977, 1988, IMAI & TAYLOR 1989). This taxon has been con-sidered a complex of morphologically similar species. IMAI & al. (1994) defined 5 different species in this complex according to their karyotypes: M. croslandi (n = 1, 2n = 2 - 4), M. imaii (2n = 6 - 8), M. banksi (2n = 9 - 10), M. haskinsorum (2n = 12 - 24), and M. pilosula s.st. (2n = 18 - 32).
Myrmecia pyriformis: See Myrmecia pilosula.
Myrmecia vindex: See Myrmecia pilosula.
Myrmica rubra: The chromosome number found for this species is 2n = 46. The populations analysed were collected in Switzerland and Japan (HAUSCHTECK 1965, IMAI 1969). Later HAUSCHTECK-JUNGEN & JUNGEN (1983) reported a 2n = 48 for Swiss populations of Myrmica laevinodis. Despite these differences, M. lae-vinodis is currently considered to be a synonym of M. rubra (BOLTON & al. 2007).
Myrmica sulcinodis: The first chromosome number given for this species was n = 24 (HAUSCHTECK-JUNGEN in CROZIER 1975), but this is probably a mistake. In HAUSCHTECK (1965) and HAUSCHTECK-JUNGEN & JUNGEN (1983) metaphase plates are presented and the haploid chromosome number is n = 28.
Odontomachus latidens: Two different chromosome num-bers have been reported for this species, n = 15 in Malaysian populations (IMAI & al. 1983) and 2n = 32 in Indonesian populations (IMAI & al. 1985). No karyotypes of metaphase plates have been published.
Odontomachus rixosus: The standard karyotype is 2n = 30 and the variation in the chromosome number in this species is due to the presence of B-chromosomes (GOСI & al. 1982, IMAI & al. 1985, 1983).
Odontoponera transversa: Indian populations were found to have 2n = 46 (IMAI & al. 1984) and Indonesian populations 2n = 42 (IMAI & al. 1985). It is not possible to compare them, since the karyotype has been published only for the Indian populations.
Pachycondyla apicalis: The variable number of chromosomes (2n = 36 to 2n = 68) has been explained by the possible existence of several species under the taxon "apicalis" (DELABIE & al. 2008).
Pachycondyla astuta: Indonesian populations were reported to have 2n = 18, 22 (IMAI & al. 1985). Only the chromosome numbers are published but without comments concerning the karyotypes.
Pachycondyla rubra: GOСI & al. (1983) explained the variable chromosome number of 2n = 38, 40 as the presence of Robertsonian polymorphisms, but neither metaphase plates nor karyotypes were presented.
Pachycondyla venerae: Probably several sibling species are under this taxon (DELABIE & al. 2008) as happen with P. apicalis (see note).
Paratrechina longicornis: In IMAI et al. (1985) give n = 16, but this may be a mistake since in another two publications n = 8 appears and the karyotype is presented in one of the studies (HUNG & al. 1972, IMAI & al. 1984).
Pheidole noda: In a Japanese population, IMAI (1969) found n = 19, 2n = 38. The analysis of new populations showed a variable chromosome number of n = 17 - 20, 2n = 37 - 39. This polymorphism is due to fusion and fission processes (IMAI & KUBOTA 1975).
Pheidole pallidula: Earlier, n = 12 was reported for this species in Swiss populations (HAUSCHTECK 1961). Later, in populations collected in Switzerland, Greece, Tunisia and Spain, an n = 10 was found (HAUSCHTECK-JUNGEN & JUNGEN 1983, PALOMEQUE & al. 1987). HAUSCHTECK-JUNGEN & JUNGEN (1983) explains this difference as a consequence of a Robertsonian polymorphism. Also, numerical variations have been found in the chromosome numbers of this species due to the presence of a B-chromosome (LORITE & al. 2000) and a polymorphism due to a pericentromeric inversion (this paper).
Plagiolepis pygmaea: The standard karyotype formula of this species is n = 7M + 1SM + 1ST. However, one population had a polymorphism due to the presence of a supernumerary chromosome segment in the long arm of a metacentric chromosome (PALOMEQUE & al. 1993a). The presence of the segment changed the chromosome morphology to submetacentric.
Plagiolepis schmitzii: A polymorphism similar to the one described in Plagiolepis pygmaea (see note) has been described in this species (PALOMEQUE & al. 1993a).
Platythyrea tricuspidata: In IMAI & al. (1983) a 2n = 96 was reported but later the authors indicated that the real chromosome number of this species is 2n = 92 -94 (IMAI & al. 1990).
Podomyrma adelaidae: While 2n = 44, 49 - 51 was re-ported (IMAI & al. 1977), the standard karyotype is 2n = 44. The other karyotypes seem to be due to the pre-sence of B-chromosomes.
Pogonomyrmex imberbiculus: The authors (TABER & al. 1988) indicated that the variation in the chromosome number n = 30, 2n = 60 - 62 was due to the presence of B-chromosomes.
Polyrhachis illaudata: While n = 18 (IMAI & al. 1983) and n = 14 (IMAI & al. 1985) have been reported as the chromosome number of this species, no karyotypes of metaphase plates have been published to perform comparisons.
Ponera scabra: In this species the chromosome numbers found are n = 3 - 4, 2n = 7 (IMAI & KUBOTA 1972, CROZIER 1975, IMAI & al. 1988). Odd diploid chromosome numbers are possible due to a chromosome fusion, since they present a large metacentric chromosome that is not present in the haploid karyotypes.
Prenolepis jerdoni: A highly variable chromosome num-ber has been found in this species, with n = 16, 20, 25, 27. The standard haploid karyotype has 16 chromo-somes and the observed variation has been explained by the presence of 4 to 11 B-chromosomes (IMAI & al. 1988a).
Pseudolasius sp. 2 (IMAI & al. 1983): Probably the standard chromosome number of this species is n = 15, 2n = 30, while n = 17 and n = 19 has also been found (IMAI & al. 1983), due to the presence of B-chromosomes.
Rhytidoponera maniae: In this species, diploid chromosome numbers of 2n = 39, 44 - 48 have been found (IMAI & al. 1977). This variation in relation to the chromosome number is a consequence of Robertsonian polymorphisms in which the increase in chromosome number is accompanied by a reduction of metacentric chromosomes and an increase in acrocentric chromosomes.
Rhytidoponera metallica: In this species 2n = 22 - 24, 36 - 37, 41 - 44 and 46 have been found (CROZIER 1969, IMAI & al. 1977). This variation is due to Robertso-nian polymorphism similar to that found in R. maniae, and at least six larger chromosome pairs were found to be involved in the Robertsonian polymorphism.
Sphinctomyrmex steinheili: 2n = 45 - 46 (IMAI & al. 1977). The variation is due to Robertsonian polymorphisms.
Strumigenys godeffroyi: Malaysian populations were found to have 2n = 40 (IMAI & al. 1983) and Indonesian populations 2n = 44 (IMAI & al. 1985). No karyotypes of metaphase plates have been published.
Tapinoma erraticum: See Tapinoma nigerrimum.
Tapinoma indicum: For this species, n = 7 was first re-ported (IMAI, in CROZIER 1975) but later IMAI & al. (1984) presented a karyotype of this species with a 2n = 10.
Tapinoma nigerrimum: Currently Tapinoma nigerrimum is considered a junior synonym of T. erraticum (BOLTON & al. 2007). However, we have considered mor-phological and karyological data that indicate that T. nigerrimum is a separate species and plan to lift the taxon from synonymy (P. Lorite, T. Palomeque & A. Tinaut, unpubl.). A polymorphism by the presence of a supernumerary chromosome segment has been detected in this species (PALOMEQUE & al. 1993a). The standard haploid formula in this species is n = 5M + 2SM + 2ST. The supernumerary chromosome segment in one of the subtelocentric chromosomes changes this chromosome to submetacentric (n = 5M + 3SM + 1ST).
Technomyrmex albipes: CROZIER (1968a) found a chromosome number of n = 9, 2n = 18 in an Australian population of this species. Afterwards, however, IMAI & al. (1977) and IMAI & al. (1984) indicated that the chromosome number for this species was n = 8, 2n = 16 in populations from Australia and India. IMAI & al. (1984) considered both karyotypes to be related by chromosome rearrangements. Crozier's population presents a karyotype with seven medium-sized metacentric chromosomes, and two small chromosomes, one metacentric and the other one acrocentric. All chromosomes in the populations analysed by IMAI & al. (1984) were found to be metacentric. These authors indicate that the two small chromosomes of the n = 9 karyotype correspond to the smallest metacentric chromosome in the n = 8 karyotype. The n = 9 karyotype derived from the n = 8 due to a centric fission and a peri-centromeric inversion.
Temnothorax albipennis: In this species, several chromo-some rearrangements have been noted, such as centric fissions and fusions, and pericentromeric inversions (FI-SCHER 1987).
Temnothorax corticalis: Although an n = 9 was initially reported (HAUSCHTECK-JUNGEN, in CROZIER 1975), later FISCHER (1987) observed that the chromosome number of this species was n = 12.
Temnothorax curvispinosus: The standard karyotype of the species is n = 23. FISCHER (1987) found the pre-sence of B-chromosomes with numbers ranging between 1 and 14.
Temnothorax nigriceps: A polymorphism by the presence of a one B-chromosome has been found in this species (FISCHER 1987).
Temnothorax rugatulus: The variation 2n = 26 - 27 (TA-BER & COKENDOLPHER 1988) found in the chromosome number is possibly due to the presence of B-chromosomes.
Temnothorax spinosior: The standard chromosome number of this species is n = 12, 2n = 24 (IMAI 1966). Later IMAI (1974) detected in this species a polymorphism due to the presence of B-chromosomes. The number of B-chromosomes ranged from one to 12 in males and their number varied among individuals of the same population.
Temnothorax unifasciatus: Temnothorax unifasciatus has a stable karyotype since material for at least six European countries has been karyotyped and a chromosome number of n = 9 has been consistently found, all without chromosome polymorphisms. T. rougeti is currently considered a non-valid taxon and a synonym of T. unifasciatus. However, the two taxa have different chromosome numbers and their karyotypes cannot easily be related. The material classified as T. rougeti has a haploid chromosome number of n = 8 and their karyotype formula is 6M + 2A, the first metacentric pair being two-fold larger than the other metacentric chromosomes (FISCHER 1987). However, the karyotype in T. unifasciatus is n = 9M, also with the first metacentric pair being two-fold larger than the other metacentric chromosomes and similar in size to the first pair of T. rougeti.
Tetramorium forte: PALOMEQUE & al. (1987) analysed the karyotype of the taxon Tetramorium hispanicum. In BOLTON & al. (2007) T. hispanicum is considered as a junior synonymy of T. ruginode. However, according to GUSTEN & al. (2006) T. hispanicum is a junior synonymy of T. forte.
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ЛИТЕРАТУРА
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